THREE spec{es of loons breed on Baffin Island--the Common
(Gayla iramet), the Black-throated (G. arctica), and the Red-throated
(G. stellata). Taverner (1934: 115) says that iraruer "does not seem
commonly to extend north of Hudson strait and the western main
coast [of the continent]." The latter part of this statement is prob-
ably correct, but in the southern half of Baffin Island iraruer breeds
regularly and in considerable numbers.
We noted all three of these loons repeatedly in the summer of 1953,
iraruer about the head of Frobisher Bay and at Lake Amadjuak;
arctica in Frobisher Bay, at Lake Amadjuak, and at Cape Dorchester;
stellata in all these areas and also at a large lake about 50 miles east-
northeast of Wordie Bay. Our findings fortify Taverner's (1934:
114) and Soper's (1928: 78) statements concerning the widespread
distribution and catholic taste of stellata. We did not, however,
find stellata nesting about the mouth of the Jordan River.
Soper (1946: 14) found that in southwestern Baffin Island the areas
actually inhabited by iraruer, arctica, and stellata to some extent
supplemented each other. He reported stellata as common and ira-
ruer as absent along the coast between Cape Dorset and Cape Dor-
chester in summer; iraruer as common and stellata as rare (or missing
altogether) "on all fresh waters" in the interior between Andrew
Gordon Bay and Tessikjuak and Ungmaluktuk lakes in September;
and iraruer as rare, stellata as scarce, and arctica as common, about
Bowman Bay in spring and summer. He mentioned one area of
breeding overlap in middle Baffin Island: Nettilling Lake, a vast body
of water in which iraruer and arctica both were "tolerably common"
in summer. He summarized thus: "These facts make it plainly
evident that the different species of loons may, on occasion, have well-
defined and locally separated breeding ranges."
We had hoped to find iraruer, arctica, and stellata all nesting in the
vicinity of the Royal Canadian Air Force Base (Lat. 63 ø 45' N.,
Long. 68 ø 33 W.) at which we had our headquarters, for we wanted
to see how nesting ponds, nest-sites, and relationships between nest
site and food supply differed among the three species. The only
species we actually found nesting close to the Base was stellata.
We saw no loon of any species in flightless condition as a result of
molt. Flying loons we saw as late as August 11 (stellata) and August
15 (iraruer and arctica). As early as July 29 we captured a flightless
molting Common Eider (Somateria mollissima); and on August 3
and 4 we saw many flightless molting Canada Geese (Branta canaden-
sis) and Blue Geese (Chen caerulescens). We suspect that many
loons, if not most of them, molt their remiges in the fall or winter
well south of their breeding grounds (see Sutton, 1943: 146).
COMMON Loon (Gayla iramet)
Kumlien (1879: 103) found this loon "common" and breeding in
Cumberland Sound. Soper (1928: 76) found it common in August
at Nettilling Lake and learned from the Eskimos that it occurred
"sparingly" about Merchants Bay in summer. Shortt and Peters
(1942: 338) were told by the Royal Canadian Mounted Police at
Pangnirtung that birds whose skins were used in Eskimo mats there-
abouts were taken locally. Wynne-Edwards (1952: 358) saw the
species in Frobisher Bay, but not at Clyde Inlet. We find no Baffin
Island records for districts north of Nettilling Lake and Merchants
Bay.
So frequently did we see a Common Loon in salt water just off the
mouth of the Sylvia Grinnell, or flying up that river, that we felt sure
at least one pair nested not very far north or northeast of the Base.
On one occasion a wild laughing cry called our attention to the flying
bird. The latest date on which we saw iraruer in this area was August 1.
On June 28, Eskimos brought to Robert Van Norman, of the Royal
Canadian Mounted Police, the fresh flat skin (minus bill, wings, and
feet) of a Common Loon recently shot just offshore from their village
near the Base.
On July 13, near the mouth of the Jordan River, we saw a Common
Loon flying low in wide circles over an Eskimo canoe not far from us.
In this same part of the bay, during stormy weather on July 20, we
chased two Common Loons with our motorboat, hoping to obtain
a specimen. We approached close enough to ascertain that they
were both adult.
On August 15, at Lake Amadjuak, we heard two Common Loons
calling loudly for a short time in the early afternoon. Presently we
saw two adults high above us. After they had circled two or three
times on rapidly beating wings, they set their wings and soared in
a graceful curve downward. They seemed to touch each other
briefly during this descent. When their wings started beating again,
the birds were about a thousand feet above the lake.
On August 8, 1955, an Eskimo named Eenutsia shot a Common
Loon (male) in full breeding feather at the head of Frobisher Bay.
The specimen was nicely prepared by Corporal Van Norman and
presented to us by him. It measures: wing (arc), 367 mm.; tail,
71; culmen, 80.5; depth of bill at base, 26.0; tarsus, 88; outer toe,
114. Average for six BaiTin Island specimens (sex undetermined)
measured by Shortt and Peters (1942: 339): wing, 356; culmen,
79.4; depth of bill, 24.6; tarsus, 87.4; outer toe, 110.
BLAC}C-T}ROATED LOON (Gayla arctica)
Taverner (1934: 115) regarded "southern Baffin" as the northern
limit of nesting for this species in eastern arctic America. Kumlien
(1879: 103) found it breeding but "not common" in Cumberland
Sound. Neither Dalgety (1936) nor Wynne-Edwards (1952) reported
it from Clyde Inlet, but Shortt and Peters (loc. cit.) saw a sin-
gle bird at Pond Inlet in September, 1938, and Soper (1928: 77)
"tentatively referred to this species" certain loons seen near Pond
Inlet in August. Soper (loc. cit.) saw the species repeatedly about
Nettilling Lake in summer and felt that the west shore was "by
reason of its low nature . . . particularly attractive and well adapted
for nesting purposes." Preference for large lakes in low country
seems to be characteristic of arctica. Rankin (1947: 50) found it
nesting in northern Scotland only in large lochs in which "sudden
influx from the surrounding country can be neutralised by the out-
flOW."
Much to our surprise we did not see arctica near the mouth of the
Sylvia Grinnell, about Tarr Inlet, or anywhere in the immediate
vicinity of the Base. On July 13, we saw an adult in the bay about
eight miles west of the Base and collected a male (GMS 11756) in
full breeding feather at the open end of a nearly ice-bound lake in
high country not far west of the Jordan River. This bird refused
to fly despite our following it about, but we failed to find its mate
or nest. It was the only loon we saw at any time in the Jordan River
district. Its eye was dark carmine, with a narrow gray ring about
the pupil (see Rankin: 1947, plate 44). The stomach was a well-
defined gizzard filled with fish remains and gravel. The testes each
measured about 20 mm. long.
At Lake Amadjuak, on August 8, we saw much of arctica. Having
ascertained that four birds well out in the lake were all adult arctica,
not two adults and their progeny, we observed two more adults flying
in. During their final descent these newcomers held their necks
strongly downward and their bills strongly upward, so the hinges
of their jaws were almost, if not quite, as low as their feet (see Fig. 1).
Of the six birds now in the water, three seemed to be paler on the head
than the others. To our surprise the flock swam slowly toward us.
When about 60 yards away they suddenly became active and noisy
FIGURI 1. Black-throated Loons coming in for a landing.
and started diving. Never were all six birds under at the same time.
Just before diving, each gave a puppy-like yelp, but the usual cry
between dives was the well-known oo-loo-lee, or a rough karr or harr,
given with a thrusting forward of the head. We did not see stellata
or iraruer anywhere in the immediate vicinity, but along another
stretch of shore we saw two stellata flying high and heard others calling
in the distance.
Near Cape Dorchester, on August 11, arctica and stellata seemed
to be about equally common, and neither showed preference, so far
as we could see, for any special sort of habitat. In a lake near which
we had heard the karr of arctica several times, we finally discovered
a half-grown, still downy chick and one of its parents. With a yelp
and noisy pattering of feet the old bird flew off. The chick dived
and we could not find it again until, some time later, both its parents
returned. One adult alighted near the spot at which we had last
seen the chick, the other in a separate pond not far away. The old
birds repeated their clear oo-loo-lee again and again, occasionally in
duet, and the chick answered with precisely the same call, in a thin,
slightly quavering voice. Witherby (1948: 120) does not mention
any such call note as this.
On August 15, in a grass-rimmed pond half a mile from the south-
east shore of Lake Amadjuak, we came upon an adult arctica and a
well-grown chick. In an arm of the lake proper we surprised a com-
pany of eight adults, all of which flew at our approach but circled
low, as if reluctant to leave.
Our male specimen of arctica measures: wing, 301 min.; tail, 55;
bill from nostril, 42; tarsus, 77. The crown, occiput and hind neck
are very pale, and the bird represents the "small" race pacifica,
though the following measurements are given by Witherby (1948:
123) for eight male G. a. arctica: wing, 280-335; tail, 50-61; bill from
nostril, 40-50; tarsus, 72-80.
RD-TIOATD Loos (Gayla stellata)
This species probably breeds throughout the whole of Baffin Island,
even in the mountainous parts. Kumlien (1879: 103) found it "very
common" in Cumberland Sound. Hantzsch (in Soper, 1928: 78)
found it "the most common of the large birds" along the south shore
of Nettilling Lake in July. Dalgety (1936: 585) found it "numerous
among the many lakes from Eglinton Fjord to Clyde Inlet" in August.
Shortt and Peters (1942: 339) saw adults and young at both Pond
Inlet and Clyde River in September. Soper (1946: 15) called stellata
"by far the commonest diver in the coastal areas of southwestern
Foxe Peninsula" and said that in southeastern Baffin Island, where
the species favored "lowlands with swampy lake surroundings,"
it nevertheless bred in "the most rugged and mountainous districts."
In this respect stellata seems to be more adaptable than arctica.
We saw stellata several times in the vicinity of the Base in the latter
half of June and in early July but did not find evidence of nesting
until July 17, when, on a lake in the middle of Hill Island, we saw a
pair of adults and two well-grown young. We failed to find stellata
at all in the pondless flat country about the mouth of the Jordan
River, and the only loon we found in the high land west of the Jordan
was arctica.
On July 30, as we were crossing the tidal flats near Davidson Point,
we saw two adult Red-throated Loons swimming in water only a
foot or so deep. Seaweed was abundant here and the loons may
have been obtaining food without diving. We expected them to
dive, swim a long way under water, and re-appear at great distance,
but instead they flew up, leaving the water with surprising ease.
On July 31, we heard two Red-throated Loons calling cocka-crah-oh,
cocka-crah-oh over and over just west of the mouth of the Sylvia
Grinnell River.
On August 3, at a pond in the middle of a great stretch of wet grass
tundra just east of Tarr Inlet, we found two adults, a half-grown
downy chick, and a nest holding one egg. The old birds flew from
the pond without a cry as we approached. The chick, which dived
well, evaded us for some time, but we finally shot it by lying in wait
well back from the shore. The nest was a muddy mass of turf on a
little island along the south shore. Well out in the pond was a rock
on which a pair of Herring Gulls (Larus argentatus) had nested. A
young gull, well feathered out but apparently unable to fly, was in
the water not far from the nest-rock. The loon chick seemed to
seek the company of this young gull.
On August 8 we saw a family of Red-throated Loons (two adults
and two young) in the middle of a large lake about 50 miles east-
northeast of Wordie Bay. That same day we saw two adult Red-
throated Loons flying high above Lake Amadjuak and heard others
calling in the far distance.
On August 11, near Cape Dorchester, we collected an adult male
and female stellata (GMS 11837, -8) at a small pond in grass tundra
bordering a low rocky ridge. We saw the pair fly in and alight and
were surprised at being able to approach them so closely. Nowhere
in that region did we hear the cries of stellata, but we heard arctica
almost constantly.
On August 15, in a small tundra pond near Lake Amadjuak, we
came upon an adult Red-throated Loon and a half-grown chick.
The birds neither dived nor tried to fly, but swam deep in the water
close to the opposite shore. At one end of the pond, in shallow water
on a muddy islet several feet out from the bank, we found the empty
nest. Not far away was part of the membrane of an egg.
Our adult male and female stellata measure: wing, 296, 272 mm.;
tail, 49, 48; bill from nostril, 39, 34; tarsus, 76, 72.
ACKNOWLEDGEMENTS
These studies were aided by a contract between the Office of Naval
Research, Department of the Navy, and the Arctic Institute of North
America N7onr-367-01. We wish to thank Robert Van Norman
and Robert Pilot, of the Royal Canadian Mounted Police, V. C.
Wynne-Edwards, of Aberdeen University, and Ross Peyton and
Brandon Halloran, of the Hudson's Bay Company, for their courtesies
and help. Especially grateful are we to L. Robert Duclos and other
officers of the Royal Canadian Air Force, for flying us to Lake Amad-
juak, Cape Dorchester, and the Wordie Bay district.
LITIRATURI CITED
DALGITY, C. r. 1936. Notes on birds observed in Greenland and Baffin Land,
June-September 1934. Ibis, 1936: 580-591.
KUMLIEN, L. 1879. Contributions to the natural history of arctic America.
U.S. Natl. Mus. Bull. 15:69-105 (Birds). Re-issued in Smithsonian Misc.
Coll., 23 (1882).
PANKIN, N. 1947. Haunts of British divers. (Collins, London and Glasgow.)
96 pp., 82 plates.
SttORTT, r. M., and H. S. PITIRS. 1942. Some recent bird records from Canada's
eastern arctic. Can. Journ. Res., I) 20: 338-348.
SoPER, J. I). 1928. A faunal investigation of southern Baffin Island. Natl. Mus.
Canada (Ottawa), Bull. 53:76-116 (Birds).
SoP, J. D. 1946. Ornithological results of the Baffin Island expeditions of
1928-1929 and 1930-1931, together with more recent records. Auk, 63: 1-24;
223-239; 418-427.
SUTTON, G.M. 1943. The wing molt of adult loons: a review of the evidence.
Wilson Bull., 55: 145-150.
TAVN, P. A. 1934. Birds of the eastern arctic. In "Canada's Eastern
Arctic": 113-128. (i)epartment of the Interior, Ottawa.)
WTHB3/4, H.F. (Editor) 1948. The handbook of British birds. Fifth impres-
sion. (Witherby and Co., London.) Vol. 4: 1-461.
WYNNe-EDWArDS, V.C. 1952. Zoology of the Baird Expedition (1950). I. The
birds observed in central and south-east Baffin Island. Auk, 69: 353-391.
Department of Zoology, University of Oklahoma, Norman, Oklahoma.