FRoM July 1958 to April 1959 I conducted a field study in eastern
Guatemala. I was accompanied throughout the nine months by my wife
Margaret and Larry L. Wolf. Dr. and Mrs. Richard R. Graber joined us
in March. We divided our time among the humid Caribbean lowlands
(Polochic Valley and lower Motagua Valley), the highlands (Sierra de
las Minas), and the arid interior, the region discussed in this report.
THE STUDY AREA
The Motague Valley and the plains about Salami, Baja Vera Paz,
constitute the arid interior covered by our work. The Rio Motague,
Guatemala's longest river, lies south of the Sierra de las Minas and flows
east to the Caribbean Sea. The upper part of the valley is the driest area
in the country, having less than 25 cm (10 inches) of rainfall per year.
Limited precipitation beginning in May initiates a short growing season.
After August rain seldom falls, and the vegetation assumes its normal
brown. Irrigation eases the farming problems of the natives.
The vegetation, controlled by aridity and sandy soil, consists of opuntias
and other cacti plus thorny scrubs of several families, notably Leguminosae.
Along the streams a more mesic flora occurs.
The extensive "Plains of Salamg" are ringed by steep ridges. The cities
of Salam and San Jer6nimo are located on the level plain at 1,000 meters
elevation. In climate the area is similar to the upper Motague, but in
vegetation grasses form the major part of the ground cover.
COLLECTING LOCALITIES
We collected specimens and took ecological and behavioral notes at 10
localities, but three fourths of the work was carried out near one station,
Usumatln.
1. Usumatldn, Zacapa. 240 meters (800 feet) elevation. Present and
past farming activities (mostly tomatoes and corn) have created cleared
areas and overgrown meadows. The original predominant growth was
scrubby woodland. This still occurs in many areas, especially on the south
slope of the Sierra de las Minas where it extends up to oak-pine forest
at 1,200 meters (4,000 feet) elevation. Along the streams a rich forest
furnishes a retreat for many species of birds in the midday heat. A limited
shore bird habitat is available. Specimens were taken at Lrsumatln during
the following periods: 9 to 19 July, 12 to 25 August, 12 September, 25
to 28 September, 22 to 28 November, 2 March, 9 March, 29 March to 3
April.
2. Quirigua, Izabal. 120 to 240 meters (400 to 800 feet). West of the
tropical village of Quirigua the oak-pine forest characteristic of the high-
lands extends down the south slope of the Sierra de las Minas into the
Motagua Valley. Here it forms the transition between the scrubby, arid
upper Motagua and the rain forests of the Caribbean lowlands. In the
valley the pines are small, and very widely spaced, dense scrub predom-
inating. Collecting dates: 6 January, 15 March.
3. Santiago, Zacapa. 180 meters (600 feet). Several kilometers west
of Quirigua the Motagua Valley takes on the almost desertlike aspect
found along most of its length. Collecting date: 22 September.
4. Zacapa, Zacapa. 240 meters (800 feet). The Motagua Valley widens
here, forming a plain 16 km across. Collecting date: 7 July.
5. Teculutdn, Zacapa. 240 meters (800 feet). This station is one of
the larger towns along the new "Route to the Atlantic," a paved road
from Guatemala City to Puerto Barrios, the major Caribbean port. Resi-
dents of Teculutn call their village the tomato capital of Guatemala.
Water for continuous fruit farming is furnished by the Teculutn River,
which flows from a deep valley in the heart of the Sierra de las Minas.
The vegetation of the area is scrubby woodland and farmland. Collecting
dates: 27 September, 11 March.
6. El Rancho, Zacapa. 300 meters (1,000 feet). The countryside is
similar to the rest of the valley. Collecting date: 28 September.
7. E1 Progreso, Guatemala. 760 meters (2,300 feet). An important
area in the ornithological history of the arid interior. The vegetation is
arid, but not as arid as at E1 Rancho or Morazn. Collecting date: 24
July.
8. Morazdn, Baja Vera Paz. 400 meters (1,200 feet). Located in a side
valley north of the Motagua, this is the driest part of the study area.
Collecting dates: 29 September, 14 November, 10 January.
9. San Jerdnimo, Baja Vera Paz. 1,000 meters (3,000 feet). Irrigated
fields and scrubby grassland surround this village at the southern edge of
the Plains of Salami. Collecting date: 10 January.
10. Salamd, Baja Vera Paz. 1,000 meters (3,000 feet). A small city
centered in the arid plains. Collecting dates: 24 February, 26 March.
PREVIOUS COLLECTING
Specimens were taken in the arid interior of Guatemala prior to 1850,
but the first work of any consequence was initiated by Osbert Sa]vin
(joined later by F. D. Godman) in 1857. Since the Motagua Valley was
on the main route used in entering or leaving the country from the east,
the birds of the arid interior received considerable attention. The results
of this work, which was continued for 40 years by professional collectors,
are recorded in the Biologia Centrali-Americana (Salvin and Godman,
1879, 1888, and 1897).
In 1906 Ned Dearborn collected at E1 Rancho, and in 1915 Rhoads
and Poole worked at Quirigua and Gualf[n. A. W. Anthony began five
years of collecting in Guatemala in 1924; in the Motagua Valley he took
specimens at Quirigua and E1 Progreso. Griscom (1932) summarized all
of these studies. Tashian (1953), in an expedition to southeastcrn
Guatemala, collected in the Motagua Valley near Teculutf[n. These early
investigators listed the following 24 species that were not collected by us.
Species names follow Eisenmann (1955), except for those included in the
A.O.U. Check-list.
Mareca americana. Baldpate. San Jer6nimo.
Buteo albicaudatus. White-tailed Hawk. San Jer6nimo, E1 Progreso.
Hypomorphnus urubitinga. Great Black Hawk. San Jer6nimo.
Harpyhaliaetus solitarius. Solitary Eagle. San Jer6nimo.
Circus cyaneus. Marsh Hawk. San Jer6nimo.
Numenius borealis. Eskimo Curlew. San Jer6nimo (now extinct).
Limnodromus griseus. Common Dowitcher. San Jer6nimo.
Erolia minutilla. Least Sandpiper. Near Quirigua.
Burhinus bistriatus. Double-striped Thick-knee. San Jer6nimo.
Speotyto cunicularia. Burrowing Owl. San Jer6nimo, GualAn, E1 Rancho.
Panyptila sanctl-hieronymi. Great Swallow-tailed Swift. San Jer6nimo.
Musclvora tyrannus. Fork-tailed Flycatcher. San Jer6nimo.
Tyrannus voclferans. Cassin's Kingbird. SalamA.
Myarchus clnerascens. Ash-throated Flycatcher. MorazAn, E1 Rancho.
Myiarchus nuttingi. Nutting's Flycatcher. E1 Rancho, E1 Progreso.
Empidonax vlrescens. Acadian Flycatcher. Near San Jer6nimo.
Camptostoma imberbe. Beardless Flycatcher. E1 Progreso.
Iridoprocne bicolor. Tree Swallow. San Jer6nimo.
Thryothorus pleurostlctus. Banded Wren. GualAn.
Salpinctes obsoletus. Rock Wren. SalamA.
Vireo flavoviridis. Yellow-green Vireo. E1 Progreso.
Parula americana. Parula Warbler. GualAn.
Dendroica coronata. Myrtle Warbler. San Jer6nimo, near Quirigua.
Pheucticus chrysopeplus. Yellow Grosbeak. E1 Rancho, E1 Progreso.
The abundance of good records from San Jer6nimo is due to the efforts
of Robert Owen, who worked under Salvin and Godman. Many of his
specimens were probably collected in the highlands, but since no elevations
were indicated, I include them all here.
DISCUSSION
Twenty-four transient species were recorded by this expedition in the
arid interior. Only one of these, Vireo bellii, was not found also in the
adjacent highlands or humid lowlands. Seventy-five resident forms were
noted. Over half of these resident birds (43 species) were recorded out-
side of the arid interior as well.
A total of 123 species (24 by previous workers) has been reported from
the arid interior of eastern Guatemala. In the adjacent Caribbean lowlands
over 300 species have been recorded. This list is restricted to material
obtained by our expedition. No attempt is made to include information
obtained, or to list specimens collected, on previous expeditions to eastern
Guatemala. Localities in italics are those where specimens were taken;
others indicate sight records only.
Butorides virescens. Green Heron. Usumatlfin. Up to 1,600 meters (5,000 feet).
Recorded in late November.
Florida caerulea. Little Blue Heron. Quirigua. Up to 260 meters (800 feet). Two
were seen on the Motagua River on 11 March.
Mycteria americana. Wood Ibis. Zacapa. Up to 260 meters (800 feet). A group
of six was seen along the Motagua River on 8 July.
Ajaia ajaja. Roseate Spoonbill. Zacapa. 200 meters. A flock of five was seen with
the Wood Ibis mentioned above.
Sarcoramphus papa. King Vulture. Usumatln. Up to 2,700 meters (8,300 feet).
Recorded occasionally.
Coragyps atratus. Black Vulture. Usumatln, Salami. Up to 1,900 meters (5,900
feet). Fairly numerous.
Cathartes aura aura. Turkey Vulture. UsumatlSn, Salami, San Jer6nimo. Up to
2,600 meters (8,000 feet). Virtually ubiquitous. More common than Coragyps
atratus. One male (collected 24 August, wing length 498 mm, tail 255 mm).
Buteo jamaicensis. Red-tailed Hawk. Morazn, Teculutn. Up to 3,100 meters
(9,500 feet). Recorded 14 November and 25 December; more common in the high
lands.
Buteo magnirostris. Roadside Hawk. Usumatln. Up to 260 meters (800 feet).
Fairly common resident in open or cultivated areas.
Buteo nitidus micrus. Gray Hawk. UsumatlSn, Quirigua. Up to 260 meters (800
feet). Less common than Buteo magnirostris. One female (collected 11 July, wing
length 259 mm, tail molting, culmen 23 mm).
Buteogallus anthracinus. Common Black Hawk. Usumatlen. Up to 260 meters
(800 feet). Uncommon resident. One female (12 August).
Herpetotheres cachinnans chapmani. Laughing Falcon. Usumatldn. Up to 260
meters (800 feet). Resident in open woodland and cultivated areas. Possibly the
most conspicuous hawk in the arid interior. One female (18 August, enlarged ovary).
Polyborus cheriway. Caracara. Usumatln, Zacapa, Teculutfin. Up to 260 meters
(800 feet). Fairly common in the upper Motagua valley in open scrubby areas.
Falco sparverius sparverius. American Sparrow Hawk. Zacapa, Usumatldn. Up to
1,600 meters (4,800 feet) (including the race F. S. tropicalis collected in the adjacent
highlands). Abundant in winter. On 11 March, while driving from Teculuffm to
Quirigua, we saw an average of almost two birds per kilometer (40 birds in 70 km).
Two females.
Colinus leucopogon. Spot-bellied Bobwhite. Usumatldn, Teculutdn, El Rancho. Up
to 260 meters (800 feet). Fairly common resident. We saw a female with several
chicks in an overgrown field on 12 August. A young bird was collected on 21 August
and an adult female from a covey of 15 on 11 March. Two females.
Charadrius collaris. Collared Plover. Usumatldn. 260 meters (800 feet). On 23
August I collected a single bird, which flushed from a small muddy pond and lit
nearby on bare ground. One male.
Tringa solitaria cinnamomea. Solitary Sandpiper. Usumatldn. 260 meters (800
feet). One male (23 August, wing length 131 mm, tail 59 mm).
Actiris macularia. Spotted Sandpiper. Usumatldn, Salamd. Up to 260 meters (800
feet). Common in winter. The earliest autumn record was on 18 August. Spotted
birds were taken in August and on 25 November, birds without spotted breasts on
30 September and 26 November. Two males, three females, one ?.
Columba livia. Rock Dove. Teculuffm. Up to 1,900 meters (5,800 feet). We re-
corded a few groups of white or largely white semiferal birds. No populations were
found established in the wild.
Zenaidura macroura marginella. Mourning Dove. Usuraatln. Up to 2,150 meters
(6,600 feet). Uncommon in winter in scrubby open woodland, more common in spring.
One male (22 November).
Zenaida asiatica asiatica. White-winged Dove. Usumatldn, E1 Rancho, Teculuffm,
Morazn, San Jer6nimo. Up to 1,600 meters (5,100 feet). Abundant resident. Speci-
mens taken in late August were molting while birds from October and November were
generally in fresh plumage. Two males, three females, one ?.
Scarda]ella inca. Inca Dove. Usumatln, Teculut/m, E1 Rancho, Morazn, San
Jer6nimo, Quirigua. Up to 1,300 meters (4,000 feet). Abundant. A nest containing
two eggs was found on 9 July. A male taken 19 July had enlarged testes. Three males.
Columbigallina passerina pallescens. Common Ground-Dove. Usuraatldn, Teculut/m,
E1 Rancho, Moraz/m, Salami, Quirigua. Up to 1,500 meters (4,500 feet). Common
in open areas. A nest was found on 12 September 1.6 meters above the ground in a
thorny tree. It contained two well-developed young. A pair with enlarged gonads
(one ovum 5 mm in length) was taken 2 March. One male, one female, one ?.
Leptotila verreauxi ]ulviventris. White-fronted Dove. Usumatldn. Up to 260
meters (800 feet). Uncommon resident in woodland along streams. A male collected
20 August was molting heavily. One male, one female.
Aratinga holochlora strenua. Green Parakeet. Progreso, Usumatlhn. Up to 750
meters (2,300 feet). Uncommon in summer in cultivated areas and scrubby woodland.
All of the specimens were collected in July and August. Four males, one female.
Amazona albi]rons nana. White-fronted Parrot. Usumatlan. Up to 330 meters
(1,000 feet). During July and August flocks of up to 20 birds were seen daily flying
into the foothills of the Sierra de las Minas in the morning and returning at dusk to
the woods along the river. A molting male was collected on 23 August. Neither this
species nor Aratinga holochlora was seen during the dry winter months. Four males.
Piaya cayana thermophila. Squirrel Cuckoo. Usumatldn, Morazn. Up to 2,100
meters (6,300 feet). Fairly common in all types of woodland. One male, one female.
Crotophaga sulcirostris sulcirostris. Groove-billed Ani. Usumatldn, San Jerdnimo,
Teculutn, Quirigua, Morazn. Up to 1,600 meters (5,000 feet). Abundant and con-
spicuous in brushy fields. Molting specimens were taken from 12 August to 24
November. A stub-tailed juvenile female was collected on 20 August. Three males,
Five females.
Morococcyx erythropygus erythropygus. Lesser Ground-Cuckoo. Usumatltn, Mora-
zion. Up to 400 meters (1,200 feet). We collected a specimen on 17 August and one
on 29 September. M. e. macrourus Griscom, described from the Motagua Valley, is
said to be paler and longer tailed than the nominate form. My two specimens are
not paler than examples of the nominate race available for comparison, and their tail
length (133 and 134 mm) is below that given by Griscom (1930) for either male or
female macrourus. Two ?.
Geococcyx velox affinis. Lesser Roadrunner. SalamiS, Usumatln, Morazn. Up to
1,000 meters (3,000 feet). Uncommon resident in open areas. The ovary of a female
taken 26 March was slightly enlarged. Two females.
Tyro alba. Barn Owl. Usumatln. 260 meters (800 feet). We saw one on 15 July.
Bubo virginianus. Great Horned Owl. Usumatln. 260 meters (800 feet). Uncom-
mon in scrubby woodland.
Glaucidium brasilianum ridgwayi. Ferruginous Pygmy-Owl. Usumatldn, Morazdn,
Quirigua. Up to 1,180 meters (3,500 feet). Common in woodland. Specimens taken
in late August and September were molting. Two males, one female.
Chordeiles acutipennis micromeris. Lesser Nighthawk. Usumatldn. 260 meters
(800 feet). We collected a single specimen 22 August on a rocky hillside. It measures:
wing 156 mm, tail 98 mm. One male.
Nyctidromus albicollis albicollis. Pauraque. El Rancho, Usumatldn. Up to 1,900
meters (5,700 feet). Common; heard calling throughout the year. Two males, one
female.
Caprimulgus ridgwayi troglodytes. Buff-collared Nightjar. Usumatldn. 260 meters
(800 feet). Specimens were taken on 13 July and 22 August on a rocky hillside
covered with scrubby vegetation. Both birds were molting. One male (wing length
143 mm), one female (wing length 152 mm).
Streptoprocne zonaris mexicana. White-collared Swift. Zacapa. Up to 2,000 meters
(6,100 feet). Two specimens were taken from a flock of birds seen at 160 meters
(500 feet) elevation on 22 September. The male specimen was molting its remiges.
One male, one ?.
Chaetura vauxi richmondi. Vaux's Swift. Usumatldn. 260 meters (800 feet). One
specimen was taken 28 September from a flock of 40 or more swifts. The bird's skull
was not ossified. A female Cypseloides rutilus was taken from the same flock. One
female.
Cypseloides rutilus brunneitorques. Chestnut-collared Swift. Usumatldn. 260 meters
(800 feet). Seen occasionally in flocks of 10 to 20 during late summer. The latest
observation was 28 September. A male collected 12 July had enlarged testes. One
male, one female.
Chlorostilbon canivetii osberti. Fork-tailed Emerald. Usumatldn. 260 meters (800
feet). Recorded on 11 March. One ?.
Amazilia rutila rutila. Cinnamon Hummingbird. Usumatldn, Quirigua. Up to 260
meters (800 feet). Uncommon resident in scrubby woodland. Four males.
Heliomaster constanti leocadiae. Plain-capped Starthroat. Usumatldn. 260 meters
(800 feet). Larry Wolf collected a male in an arid meadow on 23 November. One
male.
Trogon elegans elegans. Elegant Trogon. Usumatldn. 260 meters (800 feet). Un-
common resident in scrubby woodland. On 9 July we noted a female carrying food
into a hole five meters up in a six-meter dead stub. A male was nearby. A female taken
29 August was molting in the body. Two females.
Chloroceryle americana septentrionalis. Green Kingfisher. Usumatldn. Up to 1,600
meters (5,000 feet). Uncommon resident. One female.
Eumomota superciliosa vanrossemi. Turquoise-browed Motmot. Usumatldn. 260
meters (800 feet). Fairly common in scrubby woodland. Two molting specimens were
collected in late August. In March birds were seen flying to and from nesting holes
in sandy embankments and road cuts. Three males, one female.
Momotus mexicanus castaneiceps. Russet-crowned Motmot. Usumatldn, Morazdn,
E1 Rancho. Up to 390 meters (1,200 feet). Common in woodland. Nesting activity
was observed in March. Molting specimens were taken from 19 August to 29
November. Two males, four females.
Dryocopus lineatus similis. Lineated Woodpecker. Usumatln. Up to 1,050 meters
(3,200 feet). Fairly common. A pair collected on 9 July was molting. One male, one
female.
Melanerpes ]ormicivorus. Acorn Woodpecker. Quirigua. Up to 3,000 meters (9,000
feet). Recorded in March in open pine woodland at 200 meters (600 feet).
Centurus auri]rons santacruzi. Golden-fronted Woodpecker. Usumatldn, Salami.
Up to 1,100 meters (3,300 feet). Abundant in second-growth and open woodland. The
specimens measure: seven males, wing, 129 to 140 mm (134.8); tail, 70 to 81 mm
(77.3); one female, wing 128 mm, tail 71 mm. Seven males, one female.
Tityra semi]asciata personata. Masked Tityra. Usumatln, San Jerdnimo. Up to
2,100 meters (6,400 feet). Seen in small flocks in late summer in the Motagua Valley.
A specimen with enlarged testes was collected on 19 August. A molting bird was taken
two days later. We took another molting specimen at San Jer6nimo on 14 November.
Three males, one ?.
Muscivora ]or]icata. Sdssor-tailed Flycatcher. Zacapa, Usumatln. Up to 260
meters (800 feet). Uncommon transient and rare winter visitor.
Tyrannus verticalis. Western Kingbird. Morazn, Usumatldn. Up to 800 meters
(2,400 feet). Specimens were taken on 14 November (molting) and 25 November.
Two females.
Tyrannus melancholicus chloronotus. Tropical Kingbird. Usumatln, Quirigua,
Teculutn, Morazn, San Jer6nimo. Up to 1,600 meters (5,000 feet). Fairly common
and conspicuous resident. One male, one ?.
Myiodynastes luteiventris luteiventris. Sulphur-bellied Flycatcher. Usumatlcn,
Teculutn. Up to 260 meters (800 feet). Recorded in July and September. A speci-
men with enlarged testes was taken 10 July. Two males, one ?.
Megarynchus pitangua mexicanus. Boat-billed Flycatcher. Usumatln, San Jer6nimo,
Quirigua. Up to 800 meters (2,400 feet). Fairly common in woodland. Specimens
collected from 22 August to 23 November were molting. My specimens are much
darker below and more olive above than M.p. deserticola Griscom. This race is listed
by Griscom (1932) as occurring throughout the arid interior of Guatemala but is
probably limited to the Rio Negro Valley in the central part of the country. One
male, two females.
Pitangus sulphuratus guatimalensis. Great Kiskadee. Usumatln, Quirigua, Mora-
zn. Up to 650 meters (2,000 feet). Common and conspicuous in farmland and
second growth. A male collected 11 July had enlarged testes. Three males, one female.
Myiarchus crinitus boreus. Great Crested Flycatcher. Usumatln, Morazdn. Up to
430 meters (1,300 feet). Transient and winter visitor. Three males, one female.
Mylarchus tyrannulus cooperi. Wied's Crested Flycatcher. Usuraatldn. 260 meters
(800 feet). One female (26 November).
Contopus sordidulus veliel. Western Wood Pewee. Usuraatldn. Up to 2,400 meters
(8,700 feet). One male (18 August).
Erapidonax rainiraus. Least Flycatcher. Usuraatldn, Morazdn, San Jerdnirao,
Quirigua. Up to 800 meters (2,400 feet). Abundant in winter. The first autumn record
was on 18 August. Molting birds were taken from 25 August to 20 October. Eight
males, six females, two ?.
Tolraorayias sulphurescens cinereiceps. Yellow-olive Flycatcher. Usuraatldn. Up to
1,200 meters (3,600 feet). One male (25 November), one female (2 March).
Petrochelidon pyrrhonota pyrrhonota. Cliff Swallow. Usuraatldn, Morazn. Up to
400 meters (1,200 feet). Mixed flocks of up to 75 migrating swallows, about half of
them Barn Swallows (Hirundo rustica), the remainder mostly Cliff Swallows, were
seen in late September. The skulls of several specimens taken from these flocks were
not ossified. Four males, two females.
Hirundo rustica erythrogaster. Barn Swallow. Usuraatldn, Teculutm. Up to 330
meters (1,000 feet). Fairly common in late September. One was seen 2 March. One
male.
Tachycineta thalassina lepida. Violet-green Swallow. Usuraatldn, Morazdn, E1
Rancho. Up to 1,900 meters (5,900 feet). Common in winter. Usually seen in flocks
of up to 50 individuals. One male, two females.
Corvus corax. Common Raven. Usumatln. Up to 1,150 meters (3,500 feet). Seen
occasionally in flight on the arid south slope of the Sierra de las Minas.
Calocitta forraosa porapata. Magpie Jay. Usuraatldn, Moraztn, Quirigua. Up to
500 meters (1,500 feet). Fairly common resident in scrubby woodland. Specimens
taken 12 July and 17 August were immature; the rectrices of the latter were molting.
Three males, one ?.
Cissilopha raelanocyanea raelanocyanea. Bushy-crested Jay. Usuraatln, Salamt.
500, to 2,100 meters (1,500 to 6,500 feet). One male (24 December).
Carapylorhynchus rufinucha castaneura. Rufous-naped Wren. Usuraatln, Morazn.
Up to 1,100 meters (3,300 feet). Fairly common resident in brush, scrubby meadows,
and hedge rows. A male and female taken in mid-August had enlarged gonads. Molt-
ing specimens were collected in August and November. Five males, one female.
Troglodytes rausculus. Southern House Wren. Quirigua. Up to 3,100 meters (9,300
feet). Several were heard singing in brush in pine-oak woodland in March.
Miraus gilvus gracilis. Tropical Mockingbird. Usuraatln, Salara, Moraztn. Up to
900 meters (2,800 feet). Uncommon resident in arid meadows and scrubby woodland.
One male, two females.
Turdus grayi graff. Clay-colored Robin. Usuraatldn, San Jerdnirao, Quirigua,
Moraztn. Up to 1,000 meters (3,000 feet). Common in wooded areas along streams.
Juveniles with spotted breasts were taken on 12 August and 17 August. Molting speci-
mens were collected in October. Four males, five females, one ?.
Polioptila albiloris albiloris. White-lored Gnatcatcher. Usuraatldn, Moraztn. Up to
400 meters (1,200 feet). Fairly common resident in scrubby woodland. Males with
enlarged testes were taken on 11 March and 19 August. Four males, two females, one
?.
Vireo bellii bellii. Bell's Vireo. Usuraatldn. 260 meters (800 feet). Collected in
second growth in November and March. One male, two females.
Vireo solitarius solitarius. Solitary Vireo. Usuraatldn. Up to 2,300 meters (7,000
feet). Fairly common in winter. One female.
Vireo gilvus gilvus. Warbling Vireo. Usumatln. Up to 2,400 meters (7,300 feet).
One female (25 November).
Mnlotilta varia. Black-and-white Warbler. Usumatlrn. Up to 2,400 meters (7,300
feet). One female (22 August).
Dendrolca petechla aestiva. Yellow Warbler. Usumatlrn. Up to 1,000 meters (3,000
feet). Common in winter. Five males, one ?.
Dendroica magnolia. Magnolia Warbler. Usumatln, Quirigua. Up to 1,500 meters
(4,600 feet). Fairly common in winter. Two males.
Dendrolca townsendl. Townsend's Warbler. Morazrn. Up to 3,100 meters (9,300
feet). One male (29 September).
Oporornis tolmlel. MacGillivray's Warbler. San Jerdnlmo. 650 to 2,900 meters
(2,000 to 8,000 feet). One male (14 November).
Chamaethlypis pollocephala caninucha. Ground Chat. Usumatldn. Up to 750
meters (2,300 feet). One male (28 November).
Icterla virens virens. Yellow-breasted Chat. Usumatlrn. Up to 600 meters (1,800
feet). One ? (26 November).
Tangavlus aeneus aeneus. Bronzed Cowbird. Usumatln. Up to 260 meters (800
feet). Uncommon. A male with enlarged testes was collected on 13 July. An im-
mature male was taken 22 August. Two males.
Cassldix mexlcanus mexlcanus. Boat-tailed Grackle. Usumatltln, Teculutn. Up to
1,600 meters (4,800 feet). Fairly common; seen often in villages and cultivated areas.
A molting immature male was taken 24 August. One male, one female.
Icterus pectoralls. Spotted-breasted Oriole. Usumatltn. 1,000 meters (3,000 feet).
One was seen on 3 March in a small grove of trees.
Icterus gulars xerophilus. Black-throated Oriole. Usumatlrn, Teculutrn. 260 meters
(800 feet). Fairly common resident in arid woodlands and along streams. The speci-
mens measure: male, wing 125 mm, tail 113; female, wing 121 to 126 (123); tail 100
to 113 (105.2). One male, four females.
Icterus sclateri alticola. Streak-backed Oriole. Usumatlrn. Up to 300 meters (900
feet). Common resident in arid woodlands and meadows. Molting specimens were
obtained in August and September. Five males, four females.
Sturnella magna altlcola. Common Meadowlark. Salam& 1,000 meters (3,000 feet).
A loose flock of eight to 10 meadowlarks, from which one specimen was taken, was
seen 26 March moving northeast across the Plains of Salam in company with large
numbers of Blue Grosbeaks (Guiraca caerulea). One female.
Tanagra elegantissima. Blue-hooded Euphonia. Usumatltn. 260 meters (800 feet).
Several were seen in scrubby woodland along a stream on 11 March.
Piranga rubra rubra. Summer Tanager. Usumatltln, Quirigua. Up to 3,100 meters
(6,300 feet). Uncommon in winter. One female.
Piranga fiava figlina. Hepatic Tanager. Usumatltln. 1,150 meters (3,500 feet). A
specimen was taken in dry open woodland on 3 March. Three or four others were
seen nearby. One male (wing length 96 mm, tail 80 mm).
Pheucticus ludovicianus. Rose-breasted Grosbeak. Morazrn, Usumatltn. Up to 2,300
meters (7,000 feet). Uncommon. One female (10 January).
Guiraca caerulea chiapensls. Blue Grosbeak. Usumatldn, Salamt. Up to 1,000 meters
(3,000 feet). Resident in overgrown fields. On 26 March a scattered flock of several
dozen (race not determined) moved northeast across the Plains of Salam in company
with other migrants.
According to Stomr and Zimmerman (1959) the races Gulraca caerulea chlapensls
and G. c. laula inhabit areas to the north and south of Guatemala, respectively. The
only Guatemalan specimens listed by these workers were considered "probably inter-
mediate" between the two races. In my series of four birds the only male, an adult
taken 20 August (wing length 95 ram, tail 74, culmen 14.5), is not as bright as G. c.
lazula and seems closer to G. c. chiapensis. One male, three females.
Passerina cyanea. Indigo Bunting. Usumatlcn, San Jer6nimo. Up to 2,000 meters
(6,000 feet). Abundant winter resident in brushy meadows and second growth. Three
males, one female, 2 ?.
Passerina versicolor purpurascens. Varied Bunting. Usumatlcln. 260 meters (800
feet). Recorded in July and September in scrubby, open woodland. Males were sing-
ing on territory in July. A female taken 19 July had an enlarged ovary. We collected
a molting male 12 September. Two males, two females.
Sporophila torqueola morelleti. White-collared Seedeater. Usumatlcln. Up to 1,900
meters (5,900 feet). Abundant resident in brushy meadows. Two males, one female.
Volatinia jacarina splendens. Blue-black Grassquit. Zacapa. Up to 2,000 meters
(6,000 feet). One male (6 January).
Spinus psaltria colombianus. Lesser Goldfinch. Usumatlcln. Up to 1,600 meters
(5,000 feet). A few small flocks were seen in overgrown fields during July and August.
Three males, two ?.
Aimophila ruficauda connectens. Stripe-headed Sparrow. Usumatldn, Morazn. Up
to 300 meters (900 feet). Common in overgrown fields and hedge rows. Specimens
taken in July and November had enlarged gonads. Four males, one female, one ?.
ACKNOWLEDGMENTS
I am indebted to my wife Margaret, Larry L. Wolf, and Richard and
Jean Graber for their help in the field; to Oliver Farmen, owner of the
Zacapa Logging Company; to Dr. Jorge Ibarra, director of the Museo
Nacional de Historia Natural in Guatemala City; and to Dr. Taylor Peck,
cultural attach at the American Embassy in Guatemala.
At home, Dr. George M. Sutton, my scientific adviser, furnished con-
tinued advice throughout the study; Dr. Dean Amadon of the American
Museum of Natural History and Dr. Herbert Friedmann of the United
States National Museum permitted me to use the ornithological collections
in their care; Thomas Burleigh, Eugene Eisenmann, and Dr. Alexander
Wetmore helped with some of the more difficult identification problems.
The expedition and the subsequent work of identifying the collected
material were sponsored by a grant from the National Science Foundation.
SUMMARY
In a field study of the avifauna of the arid interior of eastern Guatemala, 75 resident and 24 migratory species were recorded. Ten stations were utilized, but most of the collecting of specimens and recording of data was done near Usumatlán. A total of 123 species is known from this region (24 reported on previous expeditions), indicating that in number of species this avifauna is one of the sparsest in Central America.
LITERATURE CITED
ESENMANN, E. 1955. The species of Middle American birds. Trans. of the Linn.
Soc., Vol. 7, 128 pp.
GRisco, L. 1932. The distribution of bird-life in Guatemala. Bull. Amer. Mus.
Nat. Hist., 64, 439 pp.
SA.VIN, O., and F. D. GODMAN. 1879.
Vol. 1, 512 pp.
SA*.VN, O., and F. D. GODAN. 1888.
Vol 2, 598 pp.
SA*.VN, O., and F. D. GODAN. 1897.
Vol 3, 510 pp.
STORER, R. W., and D. A. ZVrRAN. 1959. Variation in the Blue Grosbeak
(Gulraca caerulea) with special reference to the Mexican populations. Occ. Papers
of Mus. of Zool., Ann Arbor, Mich., No. 609.
T^smN, R.E. 1953. The birds of southeastern Guatemala. Condor, 55: 198-210.
Concord College, Athens, West Virginia.
Biologia Centrali-Americana. Zool., Aves,
Biologia Centrail-Americana. Zool., Aves,
Biologia Centrali-Americana. Zool., Aves,