Projeto Dindmica Bioldgica de Fragmentos Florestais, INPA Ecologia, C.P. 478,
Manaus, Amazonas 69011, Brazil
Recognition of the White-winged Potoo (Nyctibius
leucopterus) as distinct from the Andean Potoo (N.
maculosus; Ridgway 1912, Schulenberg et al. 1984)
highlighted the mystery surrounding the former, a
Brazilian endemic. Not reported since the type de-
scription (Wied 1821:227, 1830:311-317), N. leucopte-
rus was known only from two 19th-century museum
specimens. Here, I report the discovery of a popula-
tion of N. leucopterus and a specimen from near Ma-
naus in Amazonian Brazil (2,500 km from the type
locality in coastal eastern Brazil). I also offer the first
description of vocalizations, behavior, and diet, as
well as comment on status at ganaus, historical rec-
ords, and taxonomic status.
With their recent discovery of a new antbird from
lowland Amazonian Brazil, Lanyon et al. (1990) point-
ed out that even brief, intensive surveys in previously
unexplored parts of Amazonia can yield new bird
species. The rediscovery of N. leucopterus further em-
phasizes the importance of continued faunistic sur-
veys in the Amazon, where even the most thoroughly
studied areas like Manaus (Oren and Albuquerque
1991) still hold surprises.
On 4 October 1985, Jan P. Smith (pets. comm.) found
a roosting potoo during the day. It was perched about
20 m above the ground in a dead tree in a reserve of
the Biological Dynamics of Forest Fragments Project
(BDFFP, formerly Minimum Critical Size of Ecosys-
tems Project, 2ø30'S, 60ø0'W; for detailed description
of the BDFFP reserves, see Bierregaard and Lovejoy
1988). He identified the bird as N. leucopterus on the
basis of a "very white band on the wing" formed by
the coverts. On 14 August 1989 at 1830, after imitating
a familiar but hitherto unidentified call (described
below), I saw a small potoo fly into the crown of a
tall tree (ca. 30 m) at the edge of a BDFFP camp. Later
that night and on subsequent visits, I observed up to
two individuals. They were recognizable as N. leu-
copterus by the large, pure-white shoulder patches and
smaller size relative to the Common Potoo (N. griseus),
also present at the site.
On 19 October 1989, I visited another site about 20
km away, where I had heard the same vocalization
two years earlier. This site, also a BDFFP reserve, had
a canopy platform installed 35 m above the ground
in an emergent tree. Using alpine climbing gear (rope
Present address: Museum of Natural Science, 119
Foster Hall, Louisiana State University, Baton Rouge,
Louisiana 70803, USA.
and ascenders), I climbed to the platform at dusk and
spent the night there, whistling periodically. Finally,
at 0508 the next morning (20 October), after several
minutes of distant responses to my whistling, two
potoos approached close to me. One swooped directly
at my head and then perched in the same tree where
I stood. Locating the perched bird in the beam of my
headlamp by its bright eye shine, I collected it.
The description of the specimen is as follows: one
skin and partial skeleton, at Museu Paraense Emilio
Goeldi (MPEG 46870) in Belem, Brazil; 78.5 g; iris
yellow; mouth lining mostly pink, black distally; bill
black; feet brown; male: testis 4 x 3 ram; stomach and
contents (described below) in alcohol, at Louisiana
State University Museum of Natural Science
(LSUMNS, no. MCH 01); frozen tissue sample at
Smithsonian Institution Laboratory of Molecular
Systematics in Washington, D.C. (no. B00031).
! heard two vocalizations from N. leucopterus (re-
cording archived at Cornell University's Library of
Natural Sounds, LNS 48589, with copy at Arquivo
Sonoro Neotropical of Universidade Estadual de
Campinas, in Campinas, So Paulo). The principal
vocalization, diagnostic of the species, is a 3- to 4-s,
gradually descending, whistled glissando (Fig. 1). This
is the vocalization that I imitated to attract the birds.
Because it apparently serves in territorial advertise-
ment, I call it the song. It is similar to the song of the
Great Jacamar ([acamerops aurea), but is lower-pitched
and given at night. The song is similar to a vocaliza-
tion of the Sunbittern (Eurypyga helias), but is longer
and repeated less often (ca. 4 times/rain). The song
is given only by perched birds.
The other vocalization of N. leucopterus is a short
"bweep," sometimes repeated in rapid succession,
given while perched and in flight. It is similar to
corresponding calls of the Rufous Potoo (N. bracteatus;
unpubl. data) and those of various caprimulgids, es-
pecially the Short-tailed Nighthawk (Lurocalis semi-
torquatus), in the same region. This is apparently a
contact call, given upon arrival or in the presence of
a conspecific and was given by both individuals
whenever two were present.
The following account of behavior is based on more
than 25 encounters with this species near Manaus
from 1987 through 1991. Like many caprimulgids
(Mills 1986), N. leucopterus sang most frequently in
bright moonlight, although I did not make a rigorous
attempt to document this general observation. It was
my impression that the potoo sang only on clear nights,
especially within three or four days of the full moon.
2.0
.5
.0
0.5
Fig. 1.
I I I
I I I
3.0 3.5
1.0 1.5 2.0 2.5
Time (s)
Sohogram of Nyctibius leucopterus song (narrow-band analysis).
During this period approaching the full moon and
shortly afterwards, singing activity seemed to begin
progressively later each night, perhaps accompany-
ing the ever-later-rising moon. I could not elicit sing-
ing by imitation at other lunar phases.
This species habitually perched upright on emer-
gent snags and exposed branches in the canopy. I
never saw it lower than at canopy level (ca. 30 m),
despite efforts to coax birds down both in the forest
and at forest edges. In all observed birds, vocal imi-
tation elicited a strong approach response; however,
when I whistled from the ground, approach was lim-
ited to the nearest canopy tree or to flying back and
forth across a clearing at canopy height. By contrast,
when I whistled from the canopy, birds approached
in flight to within 1 to 2 m, and one perched as close
as7m.
Nyctibius leucopterus may use the white wing patch
in display. On several occasions, an individual that I
had whistled in showed ruffed shoulder coverts, which
effectively enlarged the white area. In dim light this
white patch was sometimes all that was visible of the
bird.
The specimen's stomach contained fragments and
entire insects of Hemiptera, Lepidoptera and Cole-
optera (including a cerambycid and three curculionid
beetles), all less than 1 cm long. No foraging was
observed.
Nyctz7ius leucopterus is probably common in the can-
opy of terra firme (upland) forest immediately north
of Manaus. Between 1987 and 1991, I and Andrew
Whittaker (pers. comm.) noted vocalizations of this
species at nine sites within the BDFFP reserves, more
or less along a 45-km linear transect. All observations
were in primary terra firme forest. Although most
observations were made from campsites (forest clear-
ings <0.5 ha near small forested streams), this prob-
ably reflects merely the great proportion of night hours
spent in camp. Jan P. Smith's (pers. comm.) sighting
was at the edge of a 10-ha forest patch, surrounded
by at least 1 km of 2-m-tall second growth, with no
stream nearby.
I noted song from two other sites, both in contig-
uous terra firme forest. One location, Reserva Flores-
tal Adolfo Ducke, lies at the northern edge of sub-
urban Manaus, approximately 50 km south of the
BDFFP sites and with a very similar avifauna (Stotz
and Bierregaard 1989). My record on 24 September
1989 represents an addition to the Ducke species list
(Willis 1977). The other site, near the ZF-2 road, lies
between Ducke and the BDFFP reserves.
The presence of N. leucopterus brings the total num-
ber of potoo species resident in the terra firme forest
north of Manaus to five, including N. griseus, N. gran-
dis (Great Potoo), N. aethereus (Long-tailed Potoo), and
N. bracteatus (see lists in Stotz and Bierregaard 1989,
Karr et al. 1990). This is the highest number of potoos
known from any site and includes all of the lowland
species. Their apparent syntopy in this area, where
from one spot I have heard four species calling on a
single night, deserves further study.
Historically, there is one specimen of N. leucopterus
(Academy of Natural Sciences of Philadelphia [ANSP]
22022) of unknown origin, labeled simply as from
"Brazil." It was undoubtedly collected in the 1800s
(M. Robbins pers. comm.). The type specimen (Amer-
ican Museum of Natural History [AMNH] 5867) is
reported as collected in the state of Bahia, eastern
Brazil, between 1816 and 1817 (Wied 1821:227, 1830:
311-317). The exact location for the type, however, is
ambiguous. In his first published reference to the
specimen, Wied (1821:227) stated that he encountered
the species in forest just outside of Conquista, the
modern city of Vit6ria da Conquista (Bokermann 1957),
on his way northeast toward Salvador. Later, he (Wied
1830:317) stated that he found the bird twice, once
near "Caravellas" and later outside of "Nazareth das
Farinhas." The former location refers to the modern
city of Caravelas (Bokermann 1957), which lies some
350 km south of Conquista. Nazareth, or Nazar, is
situated 300 km northeast of Conquista, just west of
Salvador. Both Conquista and Caravelas are variously
given as the type locality by later authors (Sclater
1866:130, Allen 1889:261, Hartert 1892:624, Ridgway
1914:587, Cory 1918:117, Pinto 1937:231, Peters 1940:
181, Greenway 1978:143).
Wied (1830:311-317) believed that he had two spec-
imens of N. leucopterus. In addition to the type, he
collected a male (AMNH 5868), which actually is a
specimen of N. griseus (Chapman 1926:274, Schulen-
berg et al. 1984; pers. observ.). Considering that Wied
passed through Caravelas a year before arriving at
Conquista and made no mention in his travel journal
(Wied 1821) of a White-winged Potoo from that re-
gion, I believe that the Caravelas bird was the N.
griseus, which Wied decided later must be conspecific
with the type N. leucopterus. The type itself presum-
ably came from somewhere along the route between
Conquista and Salvador, the expedition's last leg,
which included Nazareth (Bokermann 1957).
There is reason to suspect that Wied himself never
observed the species in the wild and that the skin
may have been brought to him by an unidentified
party. Wied (1830:317) did not state that he collected
the bird himself, but rather that he "obtained" or
"acquired" (erhielt) it. Describing its behavior, he wrote
that it "flies at dusk, often perching on the ground
on forest trails, and glides gracefully over clearings"
(Wied 1830:317). In my experience with caprimulgi-
forms, including N. leucopterus near Manaus, this is
not the behavior of potoos, which always perch up-
right on branches from which they make brief up-
ward sallies and never land on the ground. Rather,
it resembles the behavior of a caprimulgid, most prob-
ably the Pauraque (Nyctidromus albicollis), the male of
which also has conspicuous patches of white in the
wings. In the absence of any further evidence, how-
ever, we must assume that the type came from forest
in the state of Bahia, probably between Conquista and
Salvador. All three localities mentioned by Wied (1821,
1830), and most of the route between Conquista and
Salvador were probably in Atlantic coast rainforest
(Anon. 1988).
In this paper, I treat N. leucopterus as monotypic.
Peters (1940:181) and other authors have listed N.
maculosus (Ridgway 1912) as a subspecies of N. leu-
copterus, apparently based on a misinterpretation of
Chapman (1926:273). Schulenberg et al. (1984) listed
a variety of reasons to consider the two specifically
distinct, including the much smaller size of N. leu-
copterus and several plumage differences. Manaus N.
leucopterus is even smaller than the other specimens,
which it otherwise resembles closely in plumage. The
song of N. maculosus is a "loud 'raa-a'" lasting one-
half second (Schulenberg et al. 1984). This differs dra-
matically from the song of N. leucopterus and is con-
clusive evidence of the specific distinction between
the two.
The Manaus male is noticeably smaller than the
other two specimens (females) of N. leucopterus (Table
1). This difference probably is not sexual dimorphism,
because no other species of potoo (analyzed by sub-
species) shows significant sexual dimorphism in wing
chord or tail length (unpubl. data). The size difference
between the Manaus bird and the other two speci-
mens of N. leucopterus is greater than the individual
variation within any other subspecies of potoo (un-
publ. data). This suggests that Manaus N. leucopterus
represents an undescribed taxon.
TABLE 1. Comparative sizes (ram) of the three spec-
imens of Nyctibius leucopterus.
Wing
chord
(unflat-
Specimen Sex tened Tail
AMNH 5867 (type) Female 210 140
ANSP 22022 Female 211 121
MPEG 46870 (Manaus) Male 182 114
If N. leucopterus inhabits the Atlantic forest (or did
so until the recent decimation of that habitat), then
it is reasonable to expect that the Manaus bird rep-
resents a distinct taxon. Considering the disconti-
nuity of forest habitat between Amazonia and the
Atlantic coast and that Manaus N. leucopterus is ap-
parently strictly a forest bird, it is unlikely that the
species' distribution is continuous over this entire
region. Disjunct distributions including Amazonia and
the Atlantic forest of coastal Brazil are known for a
variety of forest bird taxa (Haffer 1974, 1985). Popu-
lations of the same species from the two regions are
usually considered taxonomically distinct at the sub-
species level.
A total sample of three specimens, two of uncertain
origin, is insufficient to determine the taxonomic po-
sition of the Manaus population of N. leucopterus.
Measurements of a series of individuals from Manaus
could strengthen the case for a genuine size differ-
ence between this population and the older speci-
mens. It is my hope that the information contained
here and the voice recordings will help observers to
find Nyctibius leucopterus throughout its current dis-
tribution. Being a nocturnal canopy species with an
easily confused voice, it probably has been much
overlooked.
For their help in the field, I thank Paulo Ap6stolo
Assuno, Everaldo da Costa Pereira, Albano Schulz
Neto, and especially Jairo Miranda Lopes, who swore
from the start that it was not a Sunbittern. W. E.
Magnusson courageously lent me his compressed air
rifle. R. O. Bierregaard, Jr., provided me with invalu-
able opportunities, encouragement and support
throughout my stay in Manaus and did a nice job
preparing the skin. Hugo Guimares de Mesquita
identified the stomach contents. Consultation of mu-
seum specimens was graciously facilitated by M.
LeCroy (AMNH), J. V. Reinsen, Jr. (LSUMNS), and
M. Robbins and F. Sheldon (ANSP). Michael Huth and
Alexander Sliwa translated the German. Bob Grotke
of the Cornell Library of Natural Sounds produced
the original sonogram. The manuscript benefited from
the comments of or discussions with W. P. Dunlap,
J. R. Karr, S. M. Lanyon, Rita Mesquita, J.P. O'Neill,
T. A. Parker, III, J. V. Reinsen, Jr., K. V. Rosenberg,
T. W. Sherry, D. F. Stotz, and E. O. Willis, to all of
whom I am grateful. This study was supported in part
by the World Wildlife Fund, the Instituto Nacional
de Pesquisas da AmazSnia (INPA), the Smithsonian
Institution, and the Department of Ecology, Evolu-
tion, and Organismal Biology at Tulane University;
it represents publication 87 in the Biological Dynam-
ics of Forest Fragments Project technical series.
Resumo.--Uma populao de Nycttbius leucopterus fo]
descoberta e um espcimen coletado na regio de Ma-
naus, Amazonas, 2,500 km do lugar tipo na costa atln-
tica. A espcie, distinta da espcie andina, N. macu-
losus, era anteriormente conhecida somente de dois
espcimens do sculo passado. A espcie cornurn na
floresta de terra firme ao norte de Manaus, habitando
o dossel. Incluindo N. leucopterus, esta floresta contm
cinco espcies de nictibldeo, a maior diversidade co-
nhecida em um s6 lugar. As duas vocalizaqSes, in-
cluindo um assobio descendente de 3 a 4 segundos,
so descritas pela primeira vez; a gravaqo est de-
positada no Arquivo Sonoro Neotropical da Univer-
sidade Estadual de Campinas no estado de So Paulo.
O espcimen de Manaus, depositado no Museu Pa-
raense Emilio Goeldi em Belem, menor do que os
d6is espcimens antigos, sugerindo que este repre-
senta um novo taxon. No entanto, o pequeno tamanho
da amostra no permite afirmaqo sobre sua posiqo
taxon6mica. Esta descoberta realqa a falta de conhe-
cimento da fauna da Amaz6nia, at em reas mais
estudadas como a de Manaus.
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