The Auk 112(2):521-523, 1995
Department of Biology, Caribbean Union College, P.O. Box 175,
Port of Spain, Trinidad and Tobago
As a North American who has spent three years
living in the Temperate Zone of the Neotropics (Par-
aguay), I often have wondered what the term "Neo-
tropical migrant" refers to. Although some authors
define a Neotropical migrant as any species of bird
that migrates within the Neotropical biogeographical
realm (Stangel 1992, Koford et al. 1994), the prevail-
ing view of a Neotropical migrant is a species that
breeds in North America and spends the nonbreeding
season south of the Tropic of Cancer (e.g. Hagan and
Johnston 1992a, b, Finch and Stangel 1993), a view
which excludes austral (southern) and intratropical
migrants (Levey 1994). Levey (1994) adopted a broad-
er view of the term "Neotropical migrants" by in-
cluding the latter two groups of migrants in the def-
inition. I agree with Levey's arguments that austral
and intratropical migrants need to be studied for both
scientific and conservation reasons. However, by
lureping all of these birds into the same category, we
blur the already obscure definition of what a Neo-
tropical migrant is and is not, which leaves us begging
for a more precise set of terms and definitions for the
various groups of migrants.
In the New World, there are two fundamentally
distinct systems of long-distance latitudinal migra-
tion: (1) breeding birds in temperate North America
that migrate southward to spend the winter in warm-
er climates, often in Central and South America; and
(2) breeding birds in temperate South America that
migrate northward to spend the winter in warmer
climates, but only rarely in North America. The dis-
tinctiveness of these two groups of migrants is ac-
centuated by the timing of their seasonal cycles, which
are essentially half a year apart. Biogeographers have
long agreed that most of North America (north of
central Mexico) belongs to the Nearctic realm, where-
as Central America, the Caribbean and all of South
America (south to Tierra del Fuego) belong to the
Neotropical realm (e.g. Brown and Gibson 1983; see
Fig. 1). In my mind, the two groups of migrants could
logically be referred to as: (1) "Nearctic migrants,"
which breed in temperate North America and migrate
southward; and (2) "Neotropical migrants," which
breed in temperate South America and migrate north-
ward. But instead, North Americans have persisted
in selectively applying the term "Neotropical mi-
grants" to the species breeding on the North Amer-
ican continent and wintering to the south of the Tro-
pic of Cancer, which all but precludes the use of the
term for migrant species breeding in temperate South
America. The use of this term in this context is so
pervasive that it frequently graces the titles of articles
in the major North American ornithological journals,
as well as the titles of two recently published sym-
posium volumes (Hagan and Johnston 1992b, Finch
and Stangel 1993). The term has been echoed re-
peatedly by conservation organizations and alliances,
such as Partners in Flight. As a consequence, workers
such as myself have been forced to use the term "aus-
tral migrants" in reference to migrant species breed-
ing in South America (e.g. Chesser 1994, Hayes et al.
1994, Hayes 1995).
As Levey (1994) and others have proposed, we could
simply refer to all of these migrants as "Neotropical
migrants," which is certainly logical because these
species all winter in the Neotropics and their ances-
tors probably evolved in the New World tropics (e.g.
Brown and Gibson 1983). But then what definitions
Fig. 1. The biogeographical realms of the world recognized by most biogeographers (based on Brown and
Gibson 1983).
should we use to distinguish between the two major
groups of migrants? The major geographical distinc-
tion between these groups is where they breed rather
than where they winter. Hagan and Johnston (1992a)
and Levey (1994) referred to Neotropical migrants
breeding in North America as "Nearctic-Neotropical
migrants," a precise and accurate term, but used the
vague term "austral migrants" for species breeding
in South America. These two terms are hardly com-
parable. The term "austral migrants" could equally
apply to migrant species that breed in Africa, Aus-
tralia or Antartica, which are also in the Southern
Hemisphere. Likewise, the term "boreal migrants"
(northern migrants) could apply to migrant species
breeding in either or both the Nearctic and Palearctic.
To avoid confusion and ambiguity in the use of
these terms, I recommend that a more precise ter-
minology be standardized. I propose that the follow-
ing set of terms and definitions be adopted by orni-
thologists:
Altitudinal migrant.--Any species of bird or popula-
tion of the species that regularly migrates from one
altitude to another on an annual basis within a
biogeographical realm.
Austral migrant.--Any species of bird or population
of the species breeding in the Southern Hemi-
sphere that regularly migrates northward during
the nonbreeding season.
Australian migrant.--Any species of bird or population
of the species breeding in Australia or New Zealand
that regularly migrates northward during the non-
breeding season.
Boreal migrant.--Any species of bird or population of
the species breeding in the Northern Hemisphere
that regularly migrates southward during the non-
breeding season.
Ethiopian migrant.--Any species of bird or population
of the species breeding in Africa that regularly mi-
grates northward during the nonbreeding season.
Intratropical migrant.--Any species of bird or popu-
lation of the species that breeds in the tropics and
regularly migrates to another area of the tropics on
an annual basis.
Nearctic migrant.--Any species of bird or population
of the species breeding in North America that reg-
ularly migrates southward during the nonbreeding
season.
Neotropical migrant.--Any species of bird or popula-
tion of the species breeding in South America that
regularly migrates northward during the non-
breeding season.
Palearctic migrant.--Any species of bird or population
of the species breeding in Europe or Asia that reg-
ularly migrates southward during the nonbreeding
season.
These terms are not mutually exclusive. With this
set of definitions, boreal migrants would include both
Nearctic and Palearctic migrants, whereas austral mi-
grants would include Australian, Ethiopian, and Neo-
tropical migrants. To give an example, the Peregrine
Falcon (Falco peregrinus) as a species would fit most of
these definitions; the North American populations
would be considered Nearctic migrants as well as
boreal migrants, and the South American populations
would be considered Neotropical migrants as well as
austral migrants.
I realize that such a drastic change in the meaning
of the term "Neotropical migrant" will be resisted
initially and will take time to be accepted. However,
if the status quo continues, we will continue to be
confused by a plethora of imprecise terms. Science
demands precision and logic in both our terms and
definitions. In the case of naming different groups of
migrants, the names should be based on long-ac-
cepted terms and definitions of biogeographical realms
(e.g. Brown and Gibson 1983; Fig. 1).
Given the above definitions of Nearctic and Neo-
tropical migrants, how do we distinguish between (1)
Nearctic migrants that migrate entirely within the
Nearctic, and (2) those that migrate to the Neotropics?
The distinction between these two groups of migrants
is important for both scientific and conservation rea-
sons. I suggest that the most precise terms, awkward
as they may be, are "Nearctic-Nearctic migrants" and
"Nearctic-Neotropical migrants," respectively. An al-
ternative (but less precise) set of terms might be "tem-
perate Nearctic migrants" and "tropical Nearctic mi-
grants," which could be simplified as "temperate mi-
grants" and "tropical migrants," respectively. The
same principle of combining terms could be used for
species migrating within or between other biogeo-
graphical realms (e.g. Palearctic-Palearctic migrants,
Palearctic-Ethiopian migrants, Palearctic-Oriental mi-
grants, Palearctic-Australian migrants).
Acknowledgments.--I thank J. M. Hagan III, D. W.
Johnston, D. J. Levey, and G. D. Schnell for valuable
comments and suggestions on an earlier draft of this
commentary.
LITERATURE CITED
BROWN, J. H., AND A. C. GInSON. 1983. Biogeography.
C. V. Mosby, St. Louis, Missouri.
CHESSER, R. T. 1994. Migration in South America:
An overview of the austral system. Bird Conserv.
Int. 4:91-107.
FINCH, D. M., AND P. W. STANGEL (Eds.). 1993. Status
and management of Neotropical migratory birds.
U.S. Dep. Agric. Forest Serv. Gen. Tech. Rep. RM-
229, Fort Collins, Colorado.
HAGAN, J. M., III AND D. W. JOHNSTON. 1992a. In-
troduction. Pages 1-3 in Ecology and conserva-
tion of Neotropical migrant landbirds. Smithson-
ian Institution Press, Washington, D.C.
HAGAN, J. M., III AND D. W. JOHNSTON (Eds.). 1992b.
Ecology and conservation of Neotropical migrant
landbirds. Smithsonian Institution Press, Wash-
ington, D.C.
HAYES, F. E. 1995. Status, distribution and bioge-
ography of the birds of Paraguay. Monogr. Field
Ornithol. No. 1. American Birding Association,
Colorado Springs, Colorado.
HAYES, F. E., P. A. SCHARF, AND R. S. RIDGELY. 1994.
Austral bird migrants in Paraguay. Condor 96:
83-97.
KOFORD, R. R., J. B. DUNNING, C. A. RInIC, AND D. M.
FINCH. 1994. A glossary for avian conservation
biology. Wilson Bull. 106:121-137.
LEVEY, D.J. 1994. Why we should adopt a broader
view of Neotropical migrants. Auk 111:233-236.
STANGEL, P.W. 1992. Just what is a Neotropical mi-
grant? Partners in Flight Newsl. 2(1):29. National
Fish and Wildlife Foundation, Washington, D.C.
Received 9 August 1994, accepted 27 January 1995.