T IIE following article is a report on a series of counts made of the birds on
a small area. The birds on the selected area were counted five times each
day for a period of ten days: Information was obtained about the varia-
tion in the number of species and of individuals at the different times of day as
well as about the average number of birds per unit area. The method of study
was outlined by Dr. R. M. Strong, and the work was performed by the author
during June, 1908.
The area chosen is located in the Toucher Valley, two miles east of Prescott,
,3/4alla Walla County, Washington. The region is a high rolling prairie through
wl:ich the valley runs in a general direction from east to west (see Dice, Univ.
Calif. Publ. Zool., vol. 16, 1916, pp. 293-326). The valley at this place is about
one-half mile in width, and the hills rise abruptly on either side. On account of
the relatively scanty rainfall, trees, under natural conditions, are confined to the
ground along the small river and along a little slough. Irrigation now is prac-
ticed in the valley, and numerous orchards and groves of trees occur. The soil,
both in the valley and on the hills, is rich and supports without irrigation an
abundant growth of grasses or cereals.
The census area is nearly square, being a little over 200 yards on a side. lts'
area is 9.1 acres. It extends north from the small river, including the stream and
a strip of gravel bar. Just east of the area is a group of farm buildings, and one
small barn was located near the middle of 'the area. About one-half of the area
was an open field planted in potatoes and beardless barley; a small strip near the
river and along the west side of the area had been. allowed to grow up thickly to
trees, brush, and briers; most of the remainder was in orchard and garden. The
conditions in general are typical of those found along the valley at the presen1
time, and differ considerably from the natural state.
The native trees and shrubs are willow, wild cherry, dogwood, cottonwood,
alder, birch, thorn, and elderberry. Introduced trees and shrubs growing on
the area are apple, pear, plum, peach, apricot, cherry (in fruit), locust, hazelnut,
walnut, chestnut, and osage. The cottonwoods and locusts attain a height of 80
to 100 feet.
The method of taking the census was to pursue a definite course bck and
forth through the area, going slowly and covering it quite thoroughly, a strip at
a time. The time occupied in taking each census was about one hour. The
times of starting were: 4:30 . ., 7:30 A. ., 10:30 A. ., 2 P. ., and 5 P. r.
A field glass was used and all birds possible were identified. No bird was in-
cluded in the count unless its identification was certain, and as great accur-
acy as possible was sed in counting. Some birds move so rapidly from place
to place that an absolutely accurate count is impossible; with these, all were
counted that were seen in the space being covered at the time; and it is believed
that those missed altogether would about balance those counted twice.
At the beginning of eaeb census period the temperature in the shade of t!:e
high trees was recorded, using a common thermometer, which might not have
been entirely accurate, though it shows wql the variation in temperature at. the
different periods.
The accompanying table gives the results of the census. The htal nmnber
of species observed was thirty-five. In the table they are arranged in tile order
in which they are given in a report already published on tile birds of the region
(Dice, Auk, vol. 35, 1918, pp. 40-51, 148-161). The average number of species
seen on each trip was allout fifteen. and the average number of individuals was
Fig. 19. TIIE TOUCHET VALLI.;Y TW'O MILES EAST O1.' PRESCOTT. '3/4AI,I,A
3/4ALLA COUNTY, WASInNGTON. TIlE 3/4IE' IS TOWARD TIIE NORTII-
VEST N'I) TIIE CENSUS AREA IS O TIIF RIGlIT OF TIlE RIVER, BEGIN-
N'IN'G It SHORT I}ISTANC BEYOND TIlE BARN.
about sixty-nine. Two species, the Chipping Sparrow and the Robin, were seen
at every period of observation. Four species, tile Ruffed Grouse, Say Phoebe,
Western Tanager, and Cedar Waxwing, were each seen but once. The other
species occupy intermediate positions.
From the table it will be seen that tilere is a somewhat regular variation in
the number of individuals and species seen at the different periods. The nmnber
seen is greatest in tile middle of the morning and is slightly less in the early
morning and in tile evening. It is not believed titat the nnmber of birds on
area varied mm'b at lhe different times of day or from day to day. A few
birds undoubtedly crone to the area to feed; some Lewis Woodpeckers, at least,
came from the nearby woods to feed on the ripe cherries. Other rarer birds in
moving about tbe valley occasionally visit the area; for instance, tile Ruffed
Grouse, which ordinarily lives in the denser woods along the stream, was noted
once during this census. These normal movements, with species present in
numbers, undoubtedly cause variations in the number of individuals present on
the area, and some species would only occasionally be represented. However,
the movements on and off the area would about balance each other, and the bird
population on the area should remain about the same; for species such as the
Robin, present in large numbers, even the number of individuals would probably
remain nearly constant,
The variation in the number of birds seen at the different periods of obser-
vation during the day must be due to a different proportion of those present
being seen. It will be readily admitted that the count never included all the
birds present. Many birds are very inconspicuous unless moving, and many un-
doubtedly escaped observation by keeping out of sight in the grass or brush.
Birds in song are especially likely to be noted: The relative activity of the birds
at different times of the day is probably the explanation of the variations in the
count at the various periods.
No attempt will be made to discuss the factors which might influence the
activity of the birds at different times of the day. It may be pointed out, how-
ever, that, contrary to the condition usually found during periods of migration,
the greatest number, both of species and individuals, was seen not in the very
early hours but a little later in the morning. The number of birds seen decreases
again as the heat of the day comes on, and it seems that the activity of the birds
was influenced, at least to some extent., by the temperature.
It is believed that the average total number of birds seen gives very closely
the number of individuals ordinarily present in June on the area. Probably this
average is just a little less than the number of birds actually present. Calcu-
lated to acres, our figures give 7.552 individual birds per acre of ground. As
the conditions along the Toucher Valley are very similar for at least ten miles
in either direction, it is probable that about the same number of birds per acre
will be found all along the stream in this region. The width in the valley cov.
ered by conditions like those on the census area would probably be on the average
about one-quarter mile. This would give for this season a bird population or'
about 1200 per 'unning mile along the stream. The prairie away from the vicin-
ity of the stream has a distinctly different avifauna, and supports a decidedly
]ess number of individuals per acre.
It will be noted that nearly half the individual birds seen on the census aroa
were Robins. These birds were attracted to some extent by the ripe cherries on
the area, but they are very abundant throughout the valley, being by far the
most abundant bird along the river. A number of other species of'birds, not
found on the census area during the time of this study, occur in the valley and
many of them have been seen on the area at other times. In the more heavily
forested and more secluded parts of the valley the number of individuals of tle
more retiring species is undoubtedly greater than the number recorded in this
census. It seems certain, then, that the average number of individuals per acre
of the different species can be only approximately determined from- the results
of this census. Probably for the more common species, which were seen on nearly
every trip, the results are more dependable. It would seem probable, after allow-
ing a large margin for error, that the total number of individuals per unit area
in the valley as determined by this study, can be considered quite dependable.
AVERAGES OF BIRDS SEEN ON CENSUS AREA
PREscoTT, WASHINGTON; JUNE 11-20, 1908
4:30 7:30 10:30 2:00
a.m. a.m. a.m. p.m.
Temperature. Fahr. 43.9 56.8 65.9 67.2 66.2
5:00 Aver- Aver-
p.m. Total age age
times individ- individ.
uals uals
seen
period acre
Sandhill Crane .................................... 0.1 ..................
Killdeer .................................................. 0.8 0.5 0.4 0.5
Bob-white ................................................ 0.3 1.2 1.1 1.4
Canada Ruffed Grouse ...................... 0.1 ..................
Western Mourning Dove .................. .1.4
Sparrow Hawk ...................................... 1.7
Batchelder Woodpecker ........................
Lewis Woodpecker .............................. 3.9
Red-shafted Flicker ............................ 1.3
Eastern Kingbird ................................ 1.3
Say Phoebe ................................................
Western Wood Pewee ...................... 0.5
Magpie .................................................... 4.1
Nevada Cowbird .................................. 0.4
San Diego Red-wing ................................
Western Meadowlark ........................ 0.2
Bullock Oriole ...................................... 0.3
Brewer Blackbird .............................. 1.1
Pale Goldfinch .................................... 1.1
Western Chipping Sparrow ............ 4.6
Merrill Song Sparrow ...................... 3.9
Slate-colored Fox Sparrow .............. 0.6
Black-headed Grosbeak .................... 1.7
Lazuli Bunting .................................... 1.9
Western Tanager ....................................
Cedar Waxwing ......................................
Western Warbling Vireo .................. 0.2
Eastern Yellow Warbler .................. 1.1
Macgillivray Warbler ........................ 0.4
Long-tailed Chat ....................................
Redstart ................................................ 06
Catbird .................................................. 9.5
Red-breasted Nuthatch ..........................
Long-tailed Chickadee ...................... 0.6
Western Robin ...................................... 26.4
Average species .................................. 15.3
Average individuals ............................ 61.1
0.1 2 0.04 0.004
0.9 19 0.62 0.068
1.0 29 1.00 0.110
I 0.02 0.002
3.2 2.3 2.1 1.8 40 2.16 0.237
1.3 1.6 1.1 1.2 46 1.38 0.152
0.4 0.5 0.6 0.3 15 0.36 0.040
6.0 6.8 5.6 4.8 48 5.42 0.596
1.1 2.0 1.4 1.0 33 1.36 0.149
0.4 0.9 0.8 0.8 31 0.84 0.092
0.1 ............ 1 0.02 0.002
0.4 0.2 .... 0.5 16 0.32 0.035
4.4 6.0 5.9 3.6 44 4.80 0.527
0.1 ........ 0.1 5 0.12 0.013
0.1 0.1 .... 2 0.04 0.004
0.1 0.2 0.3 0.1 8 0.18 0.020
0.4 0.4 0.4 0.2 10 0.34 0.037
.... 0.1 0.1 .... 5 0.26 0.029
0.3 0.5 0.2 0.1 19 0.44 0.048
4.9 5.2 4.9 3.9 50 4.70 0.516
4.0 2.7 3.0 2.3 49 3.18 0.349
0.3 0.5 0.2 0.3 13 0.38 0.042
1.9 3.5 1.0 1.5 46 1.92 0.211
2.4 1.5 1.6 1.3 37 1.74 0.191
........ 0.1 .... t 0.02 0.002
0.2 ........ 1 0.04 0.004
1.3 0.6 0.7 0.5 22 0.66 0.073
2.1 1.5 1.1 1.2 37 1.40 0.153
0.6 0.4 0.4 .... 15 0.36 0.040
.... 0.1 0.1 0.1 3 0.06 0.007
0.5 0.9 0.7 0.4 17 0.62 0.068
0.6 0.3 0.2 0.3 14 0.38 0.042
......... 0.3 0.1 4 0.08 0.009
4.3 3.5 0.8 2.6 22 2.36 0.259
33.9 33.6 30.3 31.3 50 31.10 3.417
16.2 15.7 14.2 14.1 15.1 ................
77.0 77.4 65.8 62.3 ...... 68.72 7.552
University of Michigan, Ann Arbor, November 4, 1920.