The Eskimo village of Ignalook on Little Diomede Island, Alaska, is isolated, and
facilities for visitors are limited. Few ornithologists have visited the island. We are
aware of no recorded observations on birds of the island in spring other than those
given here. Summer observations, mostly of a general nature, were presented by Jaques
(Auk, 46, 1929:230, and Nat. Hist. Mag., 1929:521-529) and Jenness (Can. Geog.
Jour., 7, 1933:87-92). Brooks' observations of 1953 were available to Gabrielson and
Lincoln during preparation of the "Birds of Alaska" (1959). Much of this material,
however, was not utilized. Repetition in the present paper is avoided except in a few
instances where comparisons of observations of 1953 and 1958 appear desirable.
Little Diomede is situated in Bering Strait 20 nautical miles west of Cape Prince of
Wales and 2.25 miles east of Big Diomede (USSR). Fairway Rock lies 8 miles and
King Island 52 miles southeast of Little Diomede. The island is 2 miles long, north to
south, and 1.5 miles wide, east to west. It rises steeply to a nearly flat plateau at an
altitude of 1308 feet. Vegetation is sparse and limited to patches and crevices among
the tumbled granitic boulders and outcrops. Beaches, except for a few narrow stretches
of large cobbles, are nonexistent; steep rocks and cliffs extend directly into the sea.
In the course of studies of marine mammals, Brooks spent the period from April 25
to June 26, 1953, and Kenyon from May 10 to June 14, 1958, on Little Diomede or on
the sea in its vicinity. Both of us kept systematic records of birds, although Brooks did
not attempt to list all species seen. Specimens of species presumed to be unusual at this
locality were collected whenever possible in 1958.
The season of 1953 was characterized by a period of unusual warmth in early April
but cool weather later caused an unusually late spring. However, leads were open near
Little Diomede in April and May. In 1958 only a few narrow leads opened near the
island until the latter half of May when considerable areas clear of ice appeared. This
condition apparently affected the arrival dates of murres at the island; however, breed-
ing activities commenced at about the same time as in 1953. As will be noted in the
following annotated list and table 2, the arrival dates of a number of species were
nearly the same in both seasons, as were also the periods of passage of certain migrants
going from Alaska to Siberia. A total of 40 species is listed of which 13 are known
to breed on Little Diomede.
Table 1
Estimated Magnitude of Populations of Birds Known to Breed at Little Diomede
Species Minimum Maximum
Phalacrocorax pelagicus 100 200
Larus hyperboreus 100 500
Rissa tridactyla pollicaris 20,000 100,000
Uria (aalge and lornvia) 50,000 200,000
Cepphus columba 200 500
Cyclorrhynchus psittacula 10,000 20000
Aethia cristatella 100,000 500,000
Aethia puilla 500,000 1,000,000
Fratercula corniculata 5,000 10,000
Lunda cirrhata 500 1,000
Calcarius lapponicus alascensis 50 100
Plectrophenax nivalis nivalis 200 500
Most of the small passerines observed and collected in 1958, although not previ-
ously recorded from Little Diomede, might be expected to land there en route from
wintering grounds in the Old World to breeding areas in Alaska. Species wintering in
North America may find the island a convenient resting place when blown from usual
spring migration routes.
Few attempts were made to census populations. An attempt was made to estimate a
lower and upper limit for the numbers of each species thought to be present. Large
numbers are difficult to estimate, thus the order of magnitude of the populations could
be expressed as an exponent of 10. However, when the estimated upper and lower limits
are given we believe that a more concrete expression of the approximate order of mag-
nitude of each population is achieved.
Table 2
Arrival Dates of Birds at Little Diomede
Species 1953 1958 Remarks
Chen hyperborea May 22-24 May 20-June 1 Migration passed
Grus canadensis May 15-22 May 17-20 Migration passed
Rissa tridactyla pollicaris May 15 May 14 First sighted
May 21 May 20 First on cliffs
Uria (aalge and lornvia) April 27 May 16 First seen
May 3 May 20 First on cliffs
June 11 June 9 First breeding
behavior
Cyclorrhynchus psitacula May 29 May 26 First on rocks
Aethia cristatella May 22 May 20 First seen
Aethia pula May 18 May 23 First seen
ca. May 25 May 27 First on rocks
Fratercula corniculata June 2 June 2 First seen
Lunda cirrhata June 2 May 26 First seen
Nyctea scandiaca May 23 May 26 First seen
SPECIES LIST
Fulmarus glacialis rodgersii. Fulmar. Ira N. Gabrielson (in litt.) saw only two at Little Diomede,
both white phase, near Fairway Rock on July 26, 1946. In 1953 Brooks was told by the Eskimos
that the Fulmar, locally called "walrus bird" is not known to nest on Little Diomede. In 1958 Kenyon
saw none near the cliffs. All those seen at Little Diomede in 1958 were of the white phase. The first
was seen on May 29, flying over narrow leads among ice floes about 1 mile east of Little Diomede.
Other individuals were seen on June 3, 5, and 9. On June 10, 20 Fulmars circled and fed in the
channel in front of Ignalook Village. On June 13 the birds were numerous and constantly in view
over the water from the village.
PhMacrocorax pelagicus pelagicus. Pelagic Cormorant. Brooks found cormorants present on April
25, 1953, and he believes he observed two species, one larger than the other. In 1958 only pelagicus
was noted but no special search for Phalacracorax urile was made. One cormorant was seen near the
north end of the island from the aircraft before landing on May 11, 1958. The natives said that cor-
morants are present throughout the winter, when they feed in leads, particularly around the north
point of the island'. Large numbers were not seen on the cliffs until May 18. The greatest concentra-
tions were at the north and south ends of the island. While we hunted walruses from umiaks, groups
of cormorants were often seen resting on broken pressure ridges of ice floes. Croaking mating calls
were heard from the cliffs frequently on May 20.
Olor columbianus. Whistling Swan. On May 20, 1958, two flew past Little Diomede and dis-
appeared in the direction of Siberia. They flew low over leads, passing among ice floes. The bills
appeared completely black.
Branta nigrians. Black Brant. On June 3, 1958, a flock of ten was seen flying westward toward
Big Diomede. On June 9 a subadult was seen sitting on the hiilside north of the village.
Chen hyperborea. Snow Goose. In 1953 the first flocks appeared on May 22 and by the evening
of May 24 the migration had ended. On May 20, 1958, a flock of four in company with three Sandhill
Cranes, flying in line, headed directly west over the ice south of the village. On May 23 a flock of six
was closely observed resting on an ice floe as it drifted with the current past the south point of the
island. On May 29 and 30 several flocks flew westward, and on June 1 they were heard calling while
moving over the village in a westerly direction.
Anas acura. Pintafl. On May 18, 1958, an adult male circled over the village. On May 23 one pair
was seen on a lead near shore at the south end of the island. On June 9 an adult male flew north along
the shore near the village and a male in subadult plumage was shot on the hillside behind the village.
Gonads were enlarged (25 X 7 min.). ,
Anas rea. Green-winged Teal. On June 10, 1958, an adult female (n/m/a or carotinends?)
was seen resting on an ice floe near the village. It was driven away by dogs and no others were sub-
sequently seen.
Polysticta stetieri. Stener Eider. Small flocks were seen between May 25 and June 5, 1953. None
was seen in 1958. Apparently the northward migrants usually stay close to continental coast lines.
Clangula byemalls. Oldsquaw. In 1953 these ducks appeared to pass in greatest abundance near
Cape Prince of Wales. On May 14, 1958, the first pair was seen in a lead at the south point of the
island. Pairs and small flocks, headed north or east, were seen on the open water during the entire
period.
ttistrionlcus hisrionis. Harlequin Duck. The first were seen on June 2, 1958, a flock of
proximately ten, flying northward over open leads near the south end of the island. On June 5 two
pairs were seen near shore ice north of the village.
$oner/a mo///ss/n. Common Eider. This species is often taken for food by the Eskimos. On
May 16, 1958, approximately 100 were seen on open leads south of Little Diomede. Although we
saw a number of flocks while on walrus hunting trips at sea, we also spent several entire days on the
water in late May without seeing this species.
$omaeria spectabiis. King Eider. Between May 14 and June 2', 1958, flocks of from 4 to 35
were seen on several occasions flying low over the water among ice floes. Dead individuals were seen
at several Eskimo homes on May 11 and the Eskimos said that they often take them on leads in the
winter and early spring. Brooks has observed that while great numbers migrate northward near
Cape Prince of Wales in May reltively few are seen near the Diomede Islands.
Lanpronea fisheri. Spectacled Eider. This species did not appear numerous at Little Diomede
in 1958. One adult male was brought in by a hunter on May 11 and several hunters said that they
had seen a few after this date.
Grs anadensi$. Sandhill Crane. In 1953 the migration passed between May 15 and 22. Flocks
en route from Alaska to Siberia frequently passed both north and south of Little Diomede in mid-
May, 1958. During periods of low visibility, the birds could often be heard but not seen. On May 17
four flocks were seen; on May 18 a flock of 27 and another of 24 were closely observed and a number
of others were reported by Eskimos. On May 20 a flock of 62 and 3 other individuals, accompanied
by 4 Snow Geese, passed by. A number of hours was spent in the field subsequent to May 20. On
May 23 a particular effort was made to record the passing of cranes but none was seen during nine
hours of field observations and none was reported by hunters. Apparently the entire migration passed
in four days, from May 17 to 20.
Eroia apina tcifica. Dunlin. Only stray migrants were seen. On May 20, 1958, a lone bird flew
north along the edge of the ice pack and alighted on floating ice about 2 miles east of Little Diomede.
Another was seen on the ice near shore rocks at the south end of the island on May 30. A male in
breeding condition was taken on June 1 (USNM no. 465837) as it looked for food among rocks
and snow patches at the water's edge.
Phaaropus fuicarius. Red Phalarope. A few migrants stopped at Little Diomede in late May
and June. On May 29, 1958, an adult in breeding plumage, probably a female, searched for food in
a pool of melt water on shore ice. On June 6, two flew among the ice floes about 8 miles south of
Little Diomede. Between June 10 and 14, 10 to 15 birds searched energetically for food under and
near the shore ice shelf along the west shore of Little Diomede. A male in breeding condition was
taken on June lo (USNM no. 465841).
Stercorarius pornarinus. Pomarine Jaeger. On May 20, 1958, a flock of eight, in dark plumage,
was headed north in direct flight close to the water I mile east of Little Diomede. On May 29 five
were seen and on June 10 one was seen. On June 14 and 15 several were seen at Wales where they
appeared more numerous than at Little Diomede.
Larus hyperboreus. Glaucous Gull. On June 7, 1953, the Eskimos brought in about two dozen
Glaucous Gull eggs, some of which were in late stages of incubation. Although some gulls nest on
Little Diomede, the Eskimos say that the Fairway Rock colony is larger. Gulls were relatively scarce
at Little Diomede until about May 15, 1958. On June 10 Eskimos from Diomede visited Fairway
Rock and found full sets of eggs which had been laid only a few days previously.
Rissa tridactyla pollicaris. Black-legged Kittiwake. Incubation was in progress on July 26, 1946,
when Ira N. Gabrielson visited the island (in litt.). In 1953, the first kittiwakes were seen at Little
Diomede on May 15, the first landed on the cliffs on May 21, and egg laying had not begun on
June 25.
On May 11, 1958, several flocks of 20 to 100 were seen in open leads among the ice during a
flight between Nome and Little Diomede. On May 14 four were seen flying over the ice south of
Little Diomede. As leads opened near the island, the birds became numerous. Twenty were seen on
May 16 and on May 18 huge flocks of many thousands were present on open leads near the island.
On May 20 at 8:00 a.m. thousands rested on cliff ledges, many of them on the snow. By 7:00 p.m.
thousands more had come to the cliffs. Previous to May 20 not one bird had been seen on the cliffs.
By 1:00 a.m. on May 21 the cliffs were again completely deserted by kittiwakes. Until May 26 no
birds were seen visiting the cliffs and relatively small numbers were seen near the island over open
water. On May 26 several hundred birds rested on ice and open water near shore but only about
100 to 200 were on the cliffs. The cliff population gradually increased until by May 29 every available
ledge and old nest was occupied by pairs of kittiwakes. The first subadult bird was seen on June 5
and numbers had increased considerably by June 14.
The behavior of kittiwakes, in making a preliminary visit to nesting cliffs in great numbers and
then deserting them completely for five days before returning to stay, is of interest because of its
similarity to behavior patterns of other larids. For example, the Sooty Tern (Sterns ]uscata) ap-
proaches the colony site at first by night, then in daylight for several days before finally resting on
the ground intermittently at night. The periodic approach of the flock to the colony site is described
for the Herring Gull, Larus argentatus (Tinbergen, The Herring Gull's World, 19'53:44-45).
Utqa aalge and Uria lornvia. Common Murre and Thick-biiled Murre. On April 27, 1953, large
flocks (50 to 200) began passing Little Diomede heading north, and their migration continued until
about May 5. During the same period, several thousand birds accumulated in the open leads near
Little Diomede and began landing on the cliffs at the south end of the island on May 3. Breeding was
observed on June 11, and the first eggs were found on June 19.
So many of these birds were seen that it was not ascertained which species was more numerous.
Large flocks were seen on open water on May 11, 1958, during the flight from Nome to Little Dio-
mede. Their number decreased sharply on the narrow leads 2 or 3 miles south of the island. None
was seen near Little Diomede until May 16 when leads opened near the island, at which time flocks
of several hundred were present, and by May 20 many thousands were present on the water. In the
morning none rested on the cliffs, although by 7:00 p.m. many thousands were there, most of them
sitting on snow and ice. Several birds were seen fighting and in one case two birds locked in combat
fell to the surface of the shore ice where they continued to fight, their beating wings leaving a trail
for several yards in the snow. By midnight the majority of birds had left the cliffs and by 3:00 a.m.
on the morning of May 21 not one remained. Between this date and May 31 a few birds occupied
the cliffs intermittently but at 1:00 p.m. on May 31 none was seen there. They again began to occupy
ledges by midafternoon. From this time on the cliff population steadily increased and birds were
present there at all hours. Breeding behavior was observed on June 9.
Cepphus columba. Pigeon Guillemot. In 1953 small numbers were seen on April 25, but no eggs
had been found by June 25. On May 14, 1958, two were seen in a narrow lead south of Little Dio-
mede; in the course of a day at sea on May 20 approximately 20 individuals, some in pairs, were seen.
Erratic courtship flight waz observed near an open lead on May 21. The number of pairs occupying
open water south of the island increased. On May 26 six pairs were seen near shore. By May 31 sev-
eral pairs were seen flying close to island cliffs and on June 2 several pairs occupied holes in cliffs at
the south end of the island. By June 13, when open water was available all around Little Diomede,
the number of Pigeon Guillemots had increased to a hundred or more pairs.
Cyclorrhynchus psittacula. Parakeet Auklet. In 1953 this species was first seen to settle on Little
Diomede on May 29. The first flocks of five to ten individuals were seen on May 20, 1958, well off-
shore on open water among ice floes. By 7:00 p.m., 500 to 600 birds were seen near shore. By May 23
thousands flocked on open water near shore. On May 26 auklets were flying high above the island
and a few individuals landed on the highest rocky outcrops. By May 31 large numbers began to
assemble high on the rocky slopes at about 4:30 p.m. Between this date and June 4 the numbers
increased rapidly. The Parakeet Auklet appears to be the least numerous of the three species of small
alcids. During late May, the most auklets were in the air from late afternoon until the early morning
hours in spite of the fact that daylight prevailed during 24 hours. On foggy days the auklet flights
began later than on sunny days.
Aethia cristatelIa. Crested Auklet. On May 20, 1958, three were seen flying over open leads
3 miles east of Little Diomede. By May 25 flocks of from 5 to 30 birds flew high above the upper
slopes of the island in early afternoon; by late afternoon the flocks approached the rocks closely and
a few landed on the higher slopes. An assemblage of many thousands gathered on an open lead outh
of the island, then took to the air, broke into smaller flocks and circled up about the island. The
citrus-like odor of this auklet is often quite noticeable, both on the water and in nesting areas. On
May 29 the odor of a flock of about I0,000 auklets was quite strong at least half a mile downwind
while the birds were hidden from view by fog. Large flocks on the water increased daily until by
June 3 a number of flocks consisted of an estimated 5000 to I0,000 birds. As soon as a large flock
took to the air, it broke into small groups of four to ten birds and circled upward around the island.
The greatest number of birds was in the air and on the rocks at about 2:00 a.m.
Fig. I. Habitat areas at the top of Little Diomede Island on June 2, 1958. Two Wheatears fed
among rocks at the edge of the tundra patch in the foreground. Longspurs had taken ter-
ritories on the rock-strewn plateau at left, and auklets (Aethia and Cyclorrhynchus) occu-
pied the rock slides on the right. Photograph by Kenyon, June, 1958.
Aethia pusilla. Least Auklet. The first was seen on May 23, 1958, and by May 26 a few flew
about the upper slopes. At 3:00 a.m. on May 27 thousands flew over the village and landed among
the rocks. By June 10 Least Auklets were by far the most numerous birds about Little Diomede.
The heaviest flights occurred from 9:00 p.m. to about midnight. By June 8 it appeared that the
majority of available nesting areas had been taken and flocks of 500 to 600 Least Auklets, accom-
panied hy a few of the other two species, rested on snow drifts that still covered portions of the
rocky hillside. These birds appeared to be awaiting access to nesting areas. On June 10 at twelve noon
not one bird could be seen flying. By 2:00 p.m. a few were flying and the number rapidly increased
during the afternoon and evening hours.
Fratercula corniculata. Horned Puffin. The puffins are the last summer residents to reach Little
Diomede. The first puffins arrived on June 2, 1953, and egg laying had not commenced by Juxe 25.
The first puffin was also seen flying over open water on June 2 in 1958. On June 3 two were seen on
cliffs at the south end of the island and two others flying along the cliffs. On June 6 several pairs flew
over open water and along the cliffs. On June 10 many hundreds occupied cliff ledges. This species
outnumbered the Tufted Puffin to a considerable degree.
Lunda cirrhata. Tufted Puffin. On May 26, 1958, one hird made numerous close passes along the
rocky cliffs at the south point of the island. Numbers increased s.1owly until June 13 when scattered
pairs occupied cliffside ledges.
Nyctea scandiaca. Snowy Owl. In 1953, one was seen on May 23 and the Eskimos said that this
owl is more numerous in fall than spring. One adult was seen on the top of the island on May 26, 1958.
Asio fiammeus. Short-eared Owl. Two were seen flying westward past Little Diomede on May 20
and 22, 1953. The species was not observed in 1958.
Corvus corax. Raven. On April 25, 1953, three flew from Big Diomede to Little Diomede. Sub-
sequently they were seen almost daily. On May 14, 1958, one was seen flying about near open leads
south of Little Diomede gnd on June 1 a lone bird flew from Big Diomede toward the north end
of Little Diomede. The Eskimos said that two or three Ravens always wintered at the Diomedes and
that sometimes they nested on the higher outcroppings of Little Diomede. It is doubtful that any
nested there in 1958. The species is apparently present in small numbers throughout the year.
Oenanthe oenanthe oenanthe. Wheatear. On June 2, 1958, a pair of birds was flushed from among
large boulders near the edge of the flat top of Little Diomede. They were quite wary but a female
was taken (USNM no. 465838). Although this species is recorded as breeding at Wales and a specimen
was taken on Little Diomede by Dufresne in 1924 (Gabrielson and Lincoln, The Birds of Alaska,
1959:668), it is not known to breed there.
Phylloscopus borealis kennicotti. Arctic Warbler. Only one individual was seen, on June 12, 1958.
It hopped about energetically among large granite boulders as it searched for food among lichens
and patches of dead grass near the schoolhouse. The bird, an adult female, was taken (USNM no.
465842). Although this species is not previously recorded from Little Diomede, its presence there is
not surprising. It winters in tropical Asia, has been recorded from other Bering Sea islands, and
breeds in western Alaska.
Motacilla alba. White Wagtail. Single birds were seen on June 4, 10, and 13, 1958. They were
very elusive and shy. Each bird passed rather rapidly along the island shore and did not return to
an area after leaving it. An unsuccessful effort was made to collect each individual. Identifying char-
acteristics were observed through binoculars. Nesting of this species on St. Lawrence Island is recorded
(Fay and Cade, Univ. Calif. Publ. Zool., 63, 1959:129).
Motacilia fiava tschutschensis. Yellow Wagtail. On June 13, 1958, an adult female was observed
for half an hour (12:30 p.m. to 1:00 p.m.) before it was obtained (USNM no. 465844). It repeatedly
flew to an altitude of about 75 feet over a selected spot, then while descending on rapidly vibrating
wings uttered a high-pitched trill. One other was observed later the same afternoon as it flew rapidly
along the steep Diomede hillside. The breeding range of this species includes western Alaska areas
but it has not been recorded previously from Little Diomede.
Anthus cervinus. Red-throated Pipit. On June 12 and 13, 1958, several pipits were observed as
they flew low over the rocks and grassy patches on the steep lower slopes of Little Diomede. The birds
were quite wary and exact identification of the five or six pipits seen was not certain. However, two
adult males in breeding condition were collected (USNM nos. 465843 and 465845), one on June 13
and one on June 14. This species has not previously been recorded from Little Diomede but there is
one nesting record from Wales (Bailey, Condor, 34, 1932:47). A more careful exploration of Little
Diomede may show that it breeds there also. The species is represented by only three other specimens
from North America.
Acanthis hornemanni. Hoary Redpoll. One bird was seen on May 18 and another on June 13,
1958. Both were quite light colored and were thus considered to be this species. They flitted among
rocks along a steep hillside and were so persistently chased by Snow Buntings that neither could be
obtained. Although not previously recorded from Little Diomede, redpolls probably visit there .in
small numbers on migration as they do at other Bering Sea islands.
Junco hyemalis hyemalis. Slate-colored Junco. On June 5, 1958, one bird, a male in breeding
condition, was obtained (USNM no. 465840) on the rocky hillside behind the village. The bird was
chased by a White-crowned Sparrow but repeatedly returned to one area where it was taken. Two
Eskimos said they had never seen a bird like it there before. The species has been found on other
islands of the Bering Sea but not previously on Little Diomede.
Zonotrichia leucophrys gambelii. White-crowned Sparrow. On June 5, 1958, a male in breeding
condition was first heard singing from the hillside behind the village where it remained for about
12 hours before being taken (USNM no. 465839). The Eskimos said that usually they saw one or two
of these birds in the spring, but they had never found its nest on Little Diomede. This is a common
species on the mainland of Alaska and is probably blown occasionally from its usual migration route
to islands in the Bering Sea. It is not previously recorded from Little Diomede.
Calcarius lapponicus. Lapland Longspur. In 1953 only a single pair was observed. On May 27,
1958, an adult male was seen on a grassy stretch among rocks above the village. On May 29 several
males in soaring song flight were frequently visible in this area. On June 2 about ten pairs were seen
on the fiat top of Little Diomede. A nest with five fresh eggs was found on June 9 about 300 yards
north of the schoolhouse. On June 13 this nest contained six eggs and a second nest with seven partly
incubated eggs was found nearby.
Plectrophenax nivalis nivalis. Snow Bunting. In 1953 the birds arrived in the first part of April.
Eggs were found on June 1. Birds of this species were numerous along the lower slopes of Little
Diomede on May 11, 1958. Courtship behavior was apparent on May 14. On May 18 numerous males
sang and disputed territorial boundaries among rocky outcrops. Between the hours of 11:00 p.m. and
2:00 a.m. no singing was noted. Singing began about 3:00 a.m. and continued throughout the day.
On May 23 four females and one male were seen to arrive at the island from the south. On May 24
females were first seen gathering nesting material and by May 31 numerous mated pairs occupied
territories along the shore and hillside. Only one nest, with a female incubating, was found on June 10.
One male (USNM no. 465836) was taken on May 31, 1958.
United States Fish and WildliJe Service, Seattle, Washington, and Alaska Depart-
ment o] Fish and Game, Juneau, Alaska, February 1, 1960.