For more than a decade thousands of migrants have been handled
at several banding stations, primarily in the eastern United States,
and from nocturnal kills at tall, lighted structures, and airport ceilo~
meters. These facilities provide large samples useful in studying
various aspects of avian biology. This paper presents data on
relative abundance of each sex and age group, timing of migration,
weights, and the autumn migration routes of the Black-throated
Blue Warbler (Dendroica caerulescens) and the Cape May Warbler
(D. tiqrina). Particular attention is directed to the autumn migra-
tion of these species in Florida.
MATERIALS AND METI-IODS
Most of the specimens were obtained at the recently-erected (July
1969) WDBO-WFTV TV tower near Bithlo, Orange County,
Florida. The 1,500-foot facility is the tallest structure in Florida.
Further details of the towcr's supporting structures, its lighting
system, and of the surrounding area are given by Taylor and Ander-
son (1973)L In addition to the tower samples, data from 186 Black-
throated Blue and 136 Cape May varblcrs killed during spring mi-
gration at the Vertical Assembly Building (VAB), Cape Kennedy,
Brcvard County, Florida are included.
Most of the specimens were collected during the night and in the
early morning. Many individuals had brain hemorrhages or other
damaged areas to the head; these conditions indicate that death
resulted from the birds hitting solid objects.
Nondesiccated individuals, frozen for less than three months,
were vcighcd to the nearest 0.1 g on an Ohaus triple beam balance.
Sex was determined by dissection and age by skull ossification.
To make comparisons between regions, the percentage of Black-
throated Blue and Cape May warblers among the total number of
identified parulids killed or banded was used (Tables i and 2). The
types of assumptions to which this index of relative abundance is
subject have been discussed in detail by Nisbct (1970) in his study of
nigration in the Blackpoll Warbler (Dendroica striata).
RESULTS AND DISCUSSION
Autumn migration.--Fcw autumn disasters have been reported for
100 or more Black.throated Blue or Cape May warblers killed on a
single night at tall, lighted structures and at airport cellometers in
the United States and Canada. Of 67,640 parulids reported killed
during autumn migration at these man-made structures in Canada
and in the United States, 1,653 (2.4%) were Black-throated Blue
Warblers and 597 (0.9%) were Cape May Warblc,'s (Table 1). Of
On 8 June 1973 the WDBO-WFTV TV tower suddenly collapsed, apparentl5
from being overloaded.
91,344 parulids banded during autumn migration in the United
States, 3,010 (3.3%) were Black-throated Blue Warblers and 3,684
(4.0%) were Cape May Warblers (Table 2). The data in both tables
indicate that a large number of Black-throated Blue and Cape May
warblers pass through the Atlantic Coastal States in autumn migra-
TABLE 1. Occurrences of Black-throated Blue and Cape May warblers among
warblers killed in nocturnal autumnal migration.
Total Black-throated Cape May
State/Province warblers Blue Warblers Per cent Warblers Per cent
Plains
Saskatchewan 1,072 0 0.0 0 0.0
Manitoba 589 0 0.0 3 0.5
North Dakota 154 0 0.0 0 0.0
Kansas 586 3 0.5 0 0.0
Missouri 874 I 0.1 0 0.0
Great Lakes
Minnesota 496 0 0.0 0 0.0
Wisconsin 12,228 8 0.06 246 2.0
Illinois 878 0 0.0 0 0.0
Indiana 172 2 1.1 4 2.3
Michigan 647 5 0.7 15 2.3
New York (upstate) 427 28 6.5 6 1.4
Eastern Inland
Kentucky 184 0 0.0 i 0.5
Tennessee 19,827 27 0.1 25 0.1
Atlantic Coast
New Hampshire 74 I 1.3 0
New York 715 57 7.9 16
Maryland 1,311 69 5.2 29
Washington, I). C. 892 16 1.7 8
North Carolina 3,345 318 9.5 80
South Carolina 3,149 24 0.7 5
Georgia 3,098 80 2.5 16
Jacksonville, Fla. 58 13 22.0 0
Tallahassee, Fla.. 9,089 20 0.2 4
Orlando, Fla. 7,775 981 13.0 139
0.0
22
14
08
24
01
05
00
0 O4
15
Totals 67,640 1,653 2.4 597 0.9
xdata compiled from published and unpublished studies that cover primarily these
last 10 years (see Acknowledgments and Appendix).
tion. Bell (1971) postulated that both the Cape May and Black-
throated Blue warblers, among four other parulid species, use the
mountain flyway in greater numbers than the Atlantic coast as a
regular avenue to their wintering grounds. On 5 October 1971 an
unexpected large kill of 203 Black-throated Blue Warblers occurred
at the WWAY TV tower near Boiling Springs Lakes, Brunswick
County, in southeastern North Carolina (J. Carter, unpubl. report).
In recent years large numbers of both species have been killed
during spring and autumn migrations at various localities in Brevard
County, Florida (Case, et al., 1965; L. Ellis, pers. comm.). Cooke
(1904) recognized that the Black-throated Blue Warblers migrated
during the autumn in large numbers along both coasts of southern
Florida but believed that the species was rare in central Florida.
More recently, Nisbet (1970) postulated, based upon percentage
frequencies among kills of warblers in selected states, that the Cape
T.BLr 2. Occurrences of Black-throated Blue and Cape May warblers among
warblers banded while migrating in the autumnL
Total Black-throated Cape May
Banding location warblers Blue Warblers Per cent Warblers Per cent
Manomet, Mass. 1,601 15 0.9 24 1.4
(1966-1969)
Binghamton, N.Y. 580 5 0.9 78 13.0
(1970-1972)
Island Beach, N.J. 68,802 2,205 3.2 2,693 3.9
(1956-1972)
Ship Bottom, N.J. 648 38 5.8 22 3.4
(1969-1972)
Powdermill, Penn. 13,785
(1959-1971; Leberman
and Clench, 1972)
Allegheny Front Mountain,
W. Va. 4,069
(1970, 1972)
58 0.4 431 3.1
562 13.8 425 10.4
Kiptopeke Beach, Va. 1,634 126
(1970; Bagg, 1971)
Dauphin Island, Ala.
(1970; Bagg, 1971) 225 1
Totals 91,344 3,010
77
04
33
9 0.5
0 0.0
3,684 4.0
data compiled from published and unpublished studies. Individuals who sent
results from their banding stations are mentioned in the Acknowledgments.
May Warbler's autumn migration route is similar to the Blackpoll
Warbler's transatlantic flight from New England to the Lesser
Antilles and South America. However, of 7,775 parulids killed dur-
ing four consecutive autumn migration periods at the WDBO-
WFTV facility, only 10 Blackpoll Warblers have been found com-
pared with 139 Cape May Warblers. These figures are not very
suggestive that the Cape May Warbler's autumn migration route,
especially in Florida, is similar to that of the Blackpoll. Nisbet's
(1970) data for Florida were obtained from the study of nocturnal
disasters at the WCTV tower north of Tallahassee (Stoddard and
Norris, 1967). At that tower only four Cape May and two Blackpoll
warblers were found during an l 1-year period. These figures are
much lower than the 139 Cape May and 10 Blackpoll warblers col-
lected during four years covering the same autumn months at the
central Florida facility. Not only are both of these species uncom-
mon autumn migrants in the Tallahassee area, as indicated by field
observations and the WCTV studies, but also they are uncommon
autumn migrants in northwestern Florid (Weston, 1965; Sprunt,
1954). Similar data exist for the Black-throated Blue Warbler: at
the Tallahassee tower, Stoddard and Norris (1967) reported only
19 Black-throated Blue Warblers during September, October, and
November, whereas at WDBO, 981 Black-throated Blue Warblers
were found during four years covering the same autumn months at
the central Florida facility.
In addition to the Cape May Warbler, Nisbet (1970) postulated
that the Connecticut Warbler (Oporornis agilis), Mourning Warbler
(0. philadelphia), Wilson's Warbler (Wilsonia pusilla), and Nash-
ville Warbler (Vermivora ruficapilla) have a migration route similar
to that of the Blackpoll Warbler. Each of these species was repre-
sented by four or less individuals at the WCTV tower. Since none
of the four species have hit the WDBO-WFTV facility in autumn
migration, Nisbet's (1970) ideas regarding the autumn migration
routes of these four warblers are supported.
Of 151 autumn disasters totaling one or more birds at WDBO-
WFTV, 59 contained Black-throated Blue Warblers and 25 con-
tained Cape May Warblers. Extreme dates for freshly-killed Black-
throated Blues for four autumns were 10 September and 19 Novem-
ber; Cape Mays were found from 11 September to 21 October.
Since 11 September 1969, 987 Black-throated Blue Warblers have
been collected: 335 in 1969, 34:3 in 1970, 181 in 1971, and 128 in
1972. Of 981 specimens killed in the autumn, 403 were collected in
September, 571 in October, and 7 in November. The remaining six
were found between 17 March and i May. The peak of this species'
autumn migration in central Florida evidently lies between the last
week of September to about mid-October. Since September 1969,
86 species represented by 9,129 individuals have been killed during
autumn migration at WDBO-WFTV; the Black-throated Blue
ranks the second most abundant species. The disasters at the facility
are among the largest documented for this species. The single largest
kill of 228 Black-throated Blue Warblers occurred on 28-29 Sep-
tember 1970.
Autumn migration of the Cape May Warbler through central
Florida occurs over a shorter period of time and apparently involves
fewer birds compared with the autumn migration of the Black-
throated Blue Warbler. Since 11 September 1969, 145 Cape Mays
have been collected: 17 in 1969, 68 in 1970, 24 in 1971, and 36 in
1972. Of 139 individuals killed in the autumn, 101 were collected in
September nd 38 in October. The remaining six specimens were
collected between 21 April and i May. The single largest kill took
place on 28-29 September 1970 when 51 Cape May Warblers were
found.
Age ratios.--Immature and adult Black-throated Blue and Cape
May warblers evidently migrate together because individuals of
both age groups fell from the sky at the same time during a given kill.
For the Black-throated Blue Warbler all four age-sex classes ap-
parently arrive together in central Florida. From the earliest kill
totaling 14 individuals, 3 were adult males, 3 were immature males,
4 were adult females, and 4 were immature females. Data from 880
'of 981 Black-throated Blue Warblers collected at WDBO-WFTV in
autumn migration clearly indicate the predominance of adults over
iramatures (Table 3).
TxsL: 3. Seasonal casualty totals for 880 Black-throated Blue Warblers collected
at the WDBO-WFTV TV Tower, autumns 1969-1972.
Adult Immature Adult Immature
Dates males males females females Totals
1-15 September 4 5 8 8 25
16-30 September 66 98 53 83 300
1-15 October 95 28 91 34 248
16-31 October 83 62 87 68 300
1-15 November I 2 I 1 5
16-30 November 1 i 2
Totals 250 196 240 194 880
Adults (85) outnumbered immatures in 129 Cape May Warblers
aged. The data in Table 4 indicate that a differential pattern of
migration occurs for the Cape May in central Florida. Adults pre-
dominate in the September samples whereas immatures predominate
in the smaller October samples. The first immatures were found on
25 September.
In recent years, studies from banding operations and tower kills
have shown that many migratory species in eastern North America
have a higher percentage of adults migrating southward by an inland
route than by a coastal one (Barry, 1971; Heintzelman, 1972).
These records appear to exist for both the Black-throated Blue and
Cape May warblers; however, more data from various banding and
TV tower kill studies need to be analyzed and reported. Our tower
site is inland and adults of both species clearly predominate. Coastal
sites reporting a higher percentage of immatures than adults for the
TXBLE 4. Seasonal casualty totals for 127 Cape May Warblers collected at the
WDBO-WFTV TV Tower, autumns 1969-1972.
Adult Immature Adult Immature
D ares males males females females Totals
1-15 September 14 0 5 0 19
16-30 September 30 8 27 11 76
1-15 October 4 5 i 5 15
16-31 October i 9 i 6 17
Totals 49 22 34 22 127
Black-throated Blue Warbler and/or Cape May Warbler include
Island Beach, New Jersey (Murray, 1966) and Ship Bottom, New
Jersey (R. Foy, pers. comm.). One exception to the above records
has been found. At Binghamton, New York, an inland site, F.
Marsi (pers. comm.) found 75 immature and three adult Cape May
Warblers during three autumn periods of banding.
Sex ratios.--Of 893 Black-throated Blue Warblers collected at
WDBO-WFTV in the autumn, 456 were males and 437 were females.
Adult males outnumbered individuals of the other three age-sex
groups. Of 186 specimens taken from 17 April to 14 May at the
VAB facility, 117 were males and 69 were females.
In the 129 Cape May Warblers collected at WDBO-WFTV in
autumn, 72 were males and 57 were females. Adult males out-
numbered individuals of the other age-sex classes. Of 136 specimens
taken between 17 April and 14 May at the VAB, 82 were males and
54 were females.
Weights.--Both species taken during autumn migration were rated
fat to very fat. Weights of 330 Black-throated Blue and of 75 Cape
May warblers are given in Tables 5 and 6. The mean weights for
both species regardless of age or sex are heavier than those reported
by Murray and Jehl (1964) at Island Beach, New Jersey. Mean
TABL 5. Weights (in grams) of Black-throated Blue Warblers killed at
VDBO-WFTV, September, October, and November 1969-1971.
Age and Sex Number Mean +_ S.D. Range
Adult males 93 11.9 ñ 0.90 9.6-14.0
Immature males 79 12.1 ñ 0.84 10.2-13.9
Adult females 94 11.2 __ 0.82 9.6-13.8
Immature females 64 11.2 _ 0.83 9.3-13.4
TABLE 6. Weights (in grams) of Cape May Warblers killed at WDBO-WFTV,
September and October 1969-1971.
Age and Sex Number Mean +_ S.D. Range
Adult males 24 12.6 _+ 1.01 10.2-15.2
Immature males 15 12.4 ñ 0.93 10.5-14.1
Adult females 20 12.3 ñ 1.12 10.0-14.2
Immature females 16 12.2 _ 1.05 10.0-13.5
weights of 17 male (9.0 g) and five female (9.2 g) Black-throated
Blue Warblers killed at WDBO-WFTV and the VAB during spring
migTation are lower than those of individuals killed in the autumn.
Similar data exist for 21 Cape May Warblers taken at the VAB
facility in the spring: 15 males (9.9 g), 6 females (9.4 g). An over-
water flight from the wintering grounds probably accounts for the
lighter mean weights in spring compared to those of autumn birds.
In a study of Ovenbirds (Seiurus aurocapillus) killed in central
Florida, Taylor (1972) found that spring migrants had lower mean
weights than autumn migrants.
SUMMARY
The Atlantic Coastal States are traversed by large numbers of Black-throated Blue and Cape May warblers migrating in autumn. In Florida, based upon field observations and tower studies, both species are more common in peninsular Florida than in the panhandle region in both spring and autumn. Data obtained from 981 autumn migrating Black-throated Blue and 139 Cape May warblers collected at the WDBO-WFTV TV tower in central Florida are presented. These data are supplemented by studying 186 Black-throated Blue and 136 Cape May warblers killed in the spring at the Vertical Assembly Building at Cape Kennedy. For both species, adults outnumbered immatures, males outnumbered females, and spring birds had lower mean weights than autumn birds.
ACKNOWLEDGMENTS
Thanks are extended to the owners and operators of the WDBO-
WFTV facility. Special appreciation is offered to Bruce Anderson,
Bette Schardien, and other students at FTU who have helped collect
and process the birds. Specimens from the Vertical Assembly Build-
ing were kindly donated to FTU by Lon Ellis. I thank Dr. George
A. Hall (Allegheny Front Mountain), Dr. Mary H. Clench (Powder-
mill), Mrs. Frederick Marsi (Binghamton), Mrs. Roger W. Foy
(Ship Bottom), Mrs. Katherine G. Price (Island Beach), and Mrs.
Kathleen S. Anderson (Manomet) for supplying data from their
banding stations. Unpublished data from tower kills were sent to
me from H. W. Copland (Manitoba Museum of Man and Nature),
Geoffrey Rytell (Omega Navigation Station), and J. H. Carter,
(University of North Carolina at Wilmington).
LITERATURE CITED
BXGG, A.M. 1971. The changing seasons. Amer. Birds, 25: 16-23.
BXRRY, J.J. 1971. Differential fall migration. EBBA News, 34: 55-66.
BELL, R. K. 1971. A bird bander's diary. EBBA News, 34: 39-40.
CASE, L. D., H. CRUICKSHANK, A. E. ELLIS, AND W. F. WHITE. 1965. Weather
causes heavy bird mortality. Fla. Nat., 38: 29-30.
CooKr:, W. W. 1904. Distribution and migration of North American warblers.
Biol. Surv. Bull., 18.
It.:ITZELMAN, D. S. 1972. Age classes of birds nigrating southward through
central New Jersey--autumn 1970. EBBA News, 35: 239-244.
LCEgMAN, R. C., ,Nu M. H. CcH. 1972. Bird-banding at Powdermill, 1971
and ten years reviewed. Powdermill Nature Reserve, Research Report No. 30.
MURRAY, B.G., JR. 1966. Migration of age and sex classes of passefines on the
Atlantic Coast in autumn. Auk, 83: 352-360.
MuIRY, B.G., J. and J. R. JEHL, Ja. 1964. Weights of autumn migrants from
coastal New Jersey. Bird-Banding, 35: 253-263.
NrsBET, I. C. T. 1970. Autumn migration of the Blackpoll Warbler: evidence for
long flight provided by regional survey. Bird-Banding, 41: 207-240.
SPRUNT, A., JR. 1954. Florida bird life (with addendum 1963). New York,
Coward-McCann.
SomAg), H. L., SR., xN) R. A. Nougs. 1967. Bird casualties at a Leon County,
Florida TV tower: an eleven-year study. Bull. Tall Timbers Research Station,
no. 8: 1-104.
TxYLOg, W. K. 1972. Analysis of Ovenbirds killed in central Florida. Bird-
Banding, 43: 15-19.
T.3/4Log, W. K., and B. H. Au:so. 1973. Nocturnal migrants killed at a
central Florida TV towel'; autumns 1969-1971. Wilson Bull., 85: 42-51.
Wuso, F. M. 1965. A survey of the birdlife of northwestern Florida. Bull.
Tall Timbers Research Station, no 5: 1-147.
Department of Biological Sciences, Florida Technological University,
Orlando, Florida 32816.
Received 9 Mrch 1973, ccepted 26 July 1973.
APPENDIX
leferences used in this paper but not mentioned in the Literature Cited are
listed herein.
A:, K. P. 1963. Television tower mortality in the Niagar frontier during
fall, 1962. Kingbird, 13: 192-195.
A)5s, D.A. 1962. Nocturnal migrant mortality in the Carolinas, September,
1962. Chat, 26: 83-88.
Br:cgg, M., A. Brn% x) B. Br:. 1966. The 1965 fall warbler migra-
tion at Regina. Blue Jay, 24: 10-15.
Bw:g, R., :t) J. A. E,s. 1958. An analysis of migrating birds killed at a
television tower in east-central Illinois, September 1955-May 1957. Auk,
75: 400-414.
CxL)wr:,,, L. D., xram) N. L. CvIr:g. 1963. Bird mortality at television
towers near Cdillac, Michigan. Jack-Pine Warbler, 41: 80-89.
CxgTrm, J. H., III. 1972. Avian mortality at two television towers in south-
eastern North Carolina (unpubl. report).
CIxgg,A, B. R. 1954. Disaster in migration. Chat, 18: 104-105.
Exsgt, D., G. Jcso, K. Mxgrsor, .r) D. SN3/4)R. 1961. Large bird
kills at TV towers. Bluebird, 28: 9.
E), W. H., xruJ. Hxs. 1967. Bird mortality at. KOMU-TV towel',
Columbia, Missouri, Fall 1965 and 1966. Bluebird, 34: 3-6.
GA:g, A. F. 1966. Some facts learned from nocturnal migration. Migrant, 37:
27-34.
Gown, or, M.A. 1965. Bird migration collision casualties at Saskatoon. Blue Jay,
23: 15-17.
JoursoN, D.W. 1955. Mass bird mortality in Georgia, October, 1954. Oriole,
:20: 17-26.
Jousor% D. W., u T. P. Hs. 1957. Analysis of mass bird mortality in
October, 1954. Auk, 74: 447-458.
KMrg, C.A. 1958. Destruction at the TV tower. Passenger Pigeon, 20: 3-9.
1959. More TV tower destruction. Passenger Pigeon, 21: 135-142.
Krg_ C. A., S. D. Ros, u A. C. Err,. 1964. The ornithological flood
of September 18-20, 1963. Passenger Pigeon, 26: 159-172.
K;Mrr:g, C. A., D. G. Rsw:Lr% u D. W. W.rr:g. 1966. A comparison of the
species composition of two TV tower killed samples from he same night of
migration. Wilson Bull., 78: 26-30.
L.maux, F. W. 1962. Fall migration TV tower kills, 1962. BlueJay, 20: 152.
-- 1965. ilegina and Lumsden TV tower bird mortalities, 1964. Blue Jay,
23: 18-19.
L.sKr:3/4, A. R. 1966. T. V. tower casualties at Nashville; spring and fall, 1966.
Migrant, 37: 61-62.
---- 1968. Television t. ower casualties at Nashville, autumn 1967. Migrant,
39: 25-26.
-- 1969. T. V. tower casualties at Nashville in autumn 1968. Migrant, 40:
25-26.
---- 1969. Autumn 1969 T. V. tower casualties at Nashville. Migrant, 40:
79-80.
Loan, W. G. 1951. Bird fatalities at Bluff's Lodge on the Blue Ridge Parkway,
Wilkes County, N. C. Chat, 15: 15-16.
Lov:LL, H. B. 1952. Catastrophe to birds at a Louisville airport. Kentucky
Warbler, 28: 5-6.
M.aNUWAL, D. D. 1963. TV transmitter kills in South Bend Indiana, fall 1962.
Ind. Aud. Quart., 41: 49-53.
Mr:u% J. K. 1960. An Albany airport ceilometer disaster in 1956. Kingbird,
10: 170-171.
Nrao, 1. W. 1961. legina TV tower bird mortalities--1961. Blue Jay, 19:
160-164.
NOI{THr:RN Paxton.: WxLUXF: I{ESEARCH CENTEl{.. 1972. Investigations of bird
migration and bird mortality at the Omega Navigation Station, Lamoure,
North Dakota--1971 (unpubl. report).
---- 1972. Investigations of bird migration and losses associated with the
Omega Navigation St. ation, Lamoure, North Dakota--fall 1972 (unpubl.
report).
()VrmN% 1. 1936. The 1935 fall migration at the Washington Monument.
Wilson Bull., 48: 222-224.
---- 1937. The 1936 fall migration at the Washington Monument. Wilson
Bulle., 49: 118-119.
---- 1938. The 1937 fall migration at the Washington Monument. Wilson
Bull., 50: 146.
PAALr:, P. W., Arq) B. G. P.U:MALr;:. 1959. Mortality of birds at a television
tower in central Illinois. Aud. Bull., no. 111, pp. 1-4.
PAaMXLr. r;, P. W., ANn M.D. THOMPSON. 1963. A second kill of birds at a tele-
vision tower in central Illinois. Aud. Bull., no. 128, pp. 13-1,5.
Pos% W., J. 1963. Tower casualties at Aiken, South Carolina. Chat, 27: 23.
SxwYm, P.J. 1961. Bird mortality at the WENH-TV tower in Deerfield, New
Hampshire. New Hampshire Audubon Quart., 14: 4649.
SnxaP, B. 1971. Heavy mortality of migrating birds at Madison's TV towers.
Passenger Pigeon, 33: 203-204.
StasH, L. B. 1966. Bird mortality at radio aud TV towers near Winnipeg, Sep-
tember 1965. Blue Jay, 24: 172-176.
SPooa), W. R. 1949. Mortality of birds at the ceilometer of the Nashville
airport. Wilson Bull., 61: 86-90.
8arzu% T. E. 1954. The eeilometer hazard. Chat, 18: 55-56.
Touor, H. B., .iNn R.M. M:Nr:L. 1956. Studies of birds killed in nocturnal
migration. Univ. Kan. Publ. Mts. Nat. Hist., 10: 144.