During the spring seasons of 1977-1985, birders in the western Aleutians observed large numbers of Asiatic migrants whose occurrence was relatively predictable. These birds regularly arrived during fair weather, sometimes in large mixed flocks, came from the southeast, and were exhausted upon arrival. This set of observations is consistent with an expected pattern of normal transoceanic migration. Because non-passerines seemed to arrive in the early morning and passerines in the afternoon and evening, a localized departure point is suggested. A simple model of migration mechanics suggests that departure was from the south-central Kurile Islands.
303 Dunhagen Place
Cary, North Carolina 27511 USA
OBSERVACIONES DE EMIGRANTES ASIATICOS EN LAS ALEUTIANAS
DEL OESTE
Resumen.--Desde el 1977-1985 observadores de aves localizados en las Aleutianas del oeste
han notado durante la primavera la migraci6n de un gran ndmero de aves asifiticas. E1
patr6n de movimiento es predecible; llegan las aves de forma regular cuando las condiciones
climato16gicas son buenas. Estas vienen del suroeste y como consecuencia del viaje transø
ocefinico 11egan exhaustas. Dado el caso de que las aves que no son canoras parecen llegar
temprano en la mafiana, mientras que los Passeriformes durante la tarde y la noche, se
sugiere en el trabajo un punto particular de partida localizado en las Islas Kurile del sur.
Since 1977, birders have visited the western Aleutians, principally Attu
Island, in search of Asiatic species that occur during spring migration.
Compared to North America and western Europe, relatively little is
known about east Asian migration routes, largely because of political
difficulties facing researchers (McClure 1974). Thus, the body of ob-
servers on Attu has provided an opportunistic data base by which some
aspects of east Asian migration may be understood provided that two
assumptions are true: (1) the Asiatic species recorded are representative
of the regular east Asian migration rather than "hapless" storm-blown
vagrants; (2) the migratory route taken is representative of that used by
birds which more closely track the Asian coast.
102] ,. H. Tore J. Field Ornithol.
Spring 1988
q-
Vol. 59, No. 2 Asiatic Migrants.' Western Aleutians [ 103
AREA AND METHODS
From mid-May through early June, 1977-1985, observations of Asiatic
species were made on Attu Island, Alaska in the western Aleutians.
Observations were limited almost exclusively to the southeastern coast of
the island. Although the number of observers on the island varied con-
siderably among years, all the data reported (e.g., to American Birds)
were compiled by Larry Balch, Dan Gibson, Ben King, or Thede Tobish,
all experienced observers and trip leaders. The data presented are those
reported by these individuals in American Birds and made directly avail-
able to me. In addition, I visited Attu during the 1978 and 1980 seasons
and have included data from my daily field notes. Also, I have gleaned
reports from American Birds for sightings from other islands in the
western Aleutians during the study period.
RESULTS AND DISCUSSION
From mid-May through early June, 1977-1985, at least 66 Asiatic
species were recorded in the western Aleutians, mostly Attu. Of these,
42 occurred regularly, some in moderate numbers (Table 1) and 24 were
rare to accidental in occurence (Table 2). Most of these species arrived
during fair weather and occasionally in waves. For example, from 18 to
22 May 1980, I observed several large flocks of Bramblings with an
assortment of other species (e.g., Rustic Bunting, Hawfinch, Oriental
Greenfinch, Red-throated and Olive tree-pipits) arriving daily. The flocks
included approximately 20-80 or more birds. The maximum daily count
of 156+ Bramblings (Table 1) was based on the estimates of a single
observer and was probably conservative. Other examples of sizable flocks
include 22 Terek Sandpipers that I saw in 1980 and at least 25 Lanceo-
lated Warblers reported in 1984 (Larry Balch, pers. comm.).
Over the years, migrants have arrived in distinct temporal patterns.
On Attu, migrant Asiatic shorebirds and some passerine groups (e.g.,
finches and Turdus thrushes) peak during the third week of May whereas
small Muscicapid thrushes, flycatchers and warblers peak during the first
week of June (2 wks later). These data are generally comparable with
those of mainland populations (Dement'ev et al. 1966). Typically, birds
arrived on the coast in exhausted condition. Over the next few days, many
of them drifted "inland" and then disappeared (presumably due to de-
parture). Some species evidently bred. For example, Eurasian Wigeon
and Tufted Ducks were common and usually occurred in pairs and in
1980, a pair of Bramblings were behaving as if on territory.
During the 1980 season, I found that most passerines seemed to arrive
on Attu in the afternoon and evening (Table 3). For example, about 2000
on 26 May along the coast at Murder Point, Dan Gibson found an Eye-
browed Thrush that had recently arrived. In the late evening (ca. 2100)
of 29 May also at Murder Point, Ben King found a Gray Bunting that
was exhausted and behaved as if it had just arrived. On 31 May about
1700, I observed an Olive Tree-Pipit fly in from the SSE and land
TABLE 2. Occurrence of Asiatic species recorded in only one spring season from 1977 to
1985 in the western Aleutian Islands (including Attu Island).
Number of
Species Year individuals
Spot-billed Duck (Anas poecilorhyncha)
Steller's Sea-Eagle (Haliaeetus pelagicus)
Eurasian Kestrel (Falco tinnunculus)
Northern Hobby (Falco subbuteo)
Oriental Pratincole ( Glareola maldivarus)
Common Ringed Plover (Charadrius hiaticula)
Black-winged Stilt (Himantopus himantopus)
Little Stint (Calidris minuta)
Sharp-tailed Sandpiper ( Calidris acuminata)
Curlew Sandpiper (Calidrisferruginea)
Jack Snipe (Lymnocryptes minimus)
Pin-tailed Snipe (Gallinago stenura)
Black-tailed Gull (Larus crassirostris)
Common Cuckoo (Cuculus canorus)
Lanceolated Warbler (Locustella lanceolata)
Mugimaki Flycatcher (Ficedula mugimaki)
Brown Flycatcher (Muscicapa latirostris)
Red-flanked Bluetail ( Tarsiger cyanurus)
Siberian Blue Robin (Luscinia cyane)
Brown Shrike (Lanius cristatus)
Gray Bunting (Emberiza variabilis)
Pallas' Reed-Bunting (Emberiza pallad)
Common Reed-Bunting (Emberiza schoeniclus)
Eurasian Bullfinch (Pyrrhula pyrrhula)
1985 1
1980 1
1984 1
1983 1
1985 1
1982 1
1983 1
1983 1
1980
1982 2
1981 1
1984 1
1980 1
1982 1
1984 25+
1985 1
1985 1
1982 1-2
1985 1
1984 1
1980 1
1980 1
1977 a 1
1978 1
a Six observers compared to approximately 50 in all other years.
exhausted on the rocky shore. About two hours later, a Eurasian Skylark
arrived. Conversely, non-passerines appeared to arrive late at night or
early in the morning because they were not often seen arriving but were
typically found during the first "sortie" of searches in the morning (ca.
0800-000).
I developed a simple model to predict potential departure points as-
suming an average departure time of one hour after dark (Gauthreaux
1971), and range of flight speeds of 62-66 km/h for non-passerines and
40-44 km/h for passetines (Lee 1963, Pettingill 1970, Williams et al.
1978). Great circle distances for each of nine possible departure points
were calculated (Fig. 1, Table 4). For each of these points, arrival times
on Attu were calculated assuming that the migrants flew the shortest
possible route (Table 4). Clearly, variables such as wind velocity and
direction, and deviations from the "ideal" route would affect actual arrival
times. Based on the assumption of morning arrival (e.g., 0100-0900) of
non-passerines and afternoon-evening arrival (e.g., 1400-2200) of pas-
setines, the model predicts that migrants reaching Attu embarked on their
trans-Pacific flight from the southern Kurile Islands (Fig. 1, Locations
C-E).
106] M.H. Tm,e J. Field Ornithol.
Spring 1988
T^BLE 4. Predicted schedules for Asiatic migrants arriving on Attu Island, Alaska from
nine possible departure points (see Fig. 1).
Non-passerines Passerines
Flight Average Average
Location distance to flight Estimated time flight Estimated time
on Fig. 1 Attu (km) time (h) of arrival (h) time (h) of arrival (h)
A 2618 41 1527-1757 62 1115-1715
B 2279 37 1030-1348 54 0348-0900
C 1988 31 0606-0806 47 2112-0142
D 1811 28 0339-0527 43 1727-2133
E 1651 26 0115-0251 39 1345-1733
F 1514 24 2324-0054 36 1054-1424
G 1458 23 2236-2400 35 0936-1300
H 1203 19 1857-1909 29 0403-0654
I 747 12 1233-1355 18 1815-1957
The departure points predicted by the model are supported by a variety
of empirical evidence. First, the predicted overseas flight distance of less
than 2000 km is energetically feasible. Williams et al. (1978) have shown
that western Atlantic fall migrants can successfully complete a 3500 km
journey. Second, of the 42 species that occurred two or more spring seasons
(Table 1), only three (Garganey, Common Pochard and White-throated
Needletail) apparently do not breed at or north of the latitude of the
Aleutians (Flint et al. 1984). This observation is more consistent with an
hypothesis of regular migration rather than extreme vagrancy. Alerstam
and Pettersson (1977) have noted that as a migrant nears its breeding
grounds, its tolerance for course deflections and diversions decreases. The
geography of eastern Asia (Fig. 1) is such that the further north a migrant
departs from land, the greater the angle of deflection (away from the
"ideal" flight path) to reach Attu. Conversely, the greater the distance to
the breeding grounds, the less the angle of deflection to reach Attu and
the more tolerant a bird would be of such deviations. Although significant
east-west deflections in migration routes do occur (Gauthreaux, pets.
comm.), this is an unlikely explanation for the majority of migrants seen
on Attu. Aside from the large number and predictability of migrants,
birds reaching Attu from Kamchatka would landfall on the west side of
a mountainous island 67 km long. With observations concentrated in the
southeast corner, this hypothesis would not explain exhausted birds ar-
riving from the southeast.
In summary, the Asiatic species that occur on the western Aleutians
represent a portion of the regular spring migration along the eastern coast
of Asia. Many of the birds observed arrive after an overseas flight of
1600-2000 km from the south-central Kurile Islands. Presumably, after
an appropriate rest, some of these birds complete the migration to their
60 ø N
BERING SEA
ATTU
SEA OF 'd"' ß , ?
OKHOTSK STUDY
SITE
50 ø N
PACIFIC OCEAN
150 ø E 160 ø E 170 ø E
FIGURE 1. Northwestern Pacific Ocean indicating locations of the study site on Attu Island,
Alaska and nine possible departure points for migrants reaching Attu: A--Shiriya
Point, Honshu, Japan; B--Nemuro, Hokkaido, Japan; C--Iturup Island, USSR; D--
Urup Island, USSR; E--Simushir Island, USSR; F--Rasshua Island, USSR; G--
Kharimkoton Island, USSR; H--Baykovo Island, USSR; I--Kronotskiy Peninsula,
Kamchatka, USSR.
breeding grounds in Asia. Others apparently find the Aleutians suitable
for breeding and conclude their migration there.
ACKNOWLEDGMENTS
I extend my appreciation to the United States Coast Guard which maintains Attu Island
and generously allows birders to venture there annually. I also thank Larry Balch for making
Attu accessible to the general birding populace, without whom these data would not exist.
I also thank Dr. James A. Gessaman for his assistance in preparation of the manuscript.
LITERATURE CITED
ALERSTAM, T., AND S.-G. PETTERSSON. 1977. Why do migrating birds fly along coastlines?
J. Theor. Biol. 65:699-712.
DEMENT'EV, G. P., N. A. GLADKOV, E. S. PTUSHENKO, E. P. SPANGENBERG AND A.M.
SUDILOVSKAYA. 1966. Birds of the Soviet Union. Translated from Russian by A.
Birron and Z. S. Cole. 1968. Smithsonian Institution and NSF, Washington, D.C.,
Vols. V, VI.
FLINT, V. E., a. L. BOEHME, Y. V. KOSTIN, AND A. A. KUZNETSOV. 1984. A field guide
to birds of the USSR. Translated from Russian by N. Bourso-Leland. Princeton Uni-
versity Press, Princeton, New Jersey.
GAUTHREAUX, S. A., JR. 1971. A radar and direct visual study of passerinc spring migration
in southern Louisiana. Auk 88:343-365.
LEE, S. L. B. 1963. Migration in the Outer Hebrides. Ibis 105:493-515.
MCCLURE, H. E. 1974. Migration and survival of birds of Asia. U.S. Army Medical
Component, S.E.A.T.O., Thailand.
PETTINGILL, O. S., JR. 1970. Ornithology in laboratory and field. Burgess Publ., Min-
neapolis, Minnesota.
WILLIAMS, T. C., J. M. WILLIAMS, C. C. IRELAND, AND J. M. TEAL. 1978. Estimated
flight time for transatlantic autumnal migrants. Am. Birds 32:275-280.
Received 5 Nov. 1986; accepted 21 Nov. 1987.