At minimum ages of 24 yr-2 mo and 25 yr-9 mo, four Marbled Godwits (Limosa fedoa), banded as adults in 1969 and observed during 1992-1994 at Humboldt Bay, California, surpass the published North American longevity record for scolopacids. Minimum annual survival rates (87%) of Marbled Godwits based on these four birds are consistent with published estimates based on resighting of color-marked Palearctic shorebirds of similar size, but estimates exceed those based on recoveries of banded birds.
Department of Wildlife
Humboldt State University
Arcata, California 95521 USA
CUATRO INDIVIDUOS DE LIMOSA FEDOA EXCEDEN EL RICORD DE
LONGEVIDAD INFORMADO PARA ESCOLOPJ. CIDOS
Sinopsis.--A edades minimas de 24 aftos con dos meses y 25 aftos con nueve meses, cuatro
individuos de Limosafedoa, anillados como adultos en el 1969 y observados durante 1992-
1994 en la Bahia de Humboldt, California, sobrepasan el rcord de longevidad para escol-
opcidos. La tasa minima de sobrevivencia anual (87%) para esta especie, basada en las
cuatro aves indicadas en este trabajo, es consistente con estimados (basados en el reavista-
miento de aves marcadas con marbetes de colores) publicados para playeros paleogrticos de
tamafio similar. No obstante, los estimados exceden aquellos procedentes del recobro de aves
anilladas.
Recently, Marks (1992) reported the longevity record (minimum age
23 yr-10 mo) for North American scolopacids based on a Bristle-thighed
Curlew (Numenius tahitiensis) collected at Laysan Island in the North-
western Hawaiian Islands. Here, we report independent sightings of four
Marbled Godwits (Limosafedoa) from Humboldt Bay, California (40ø5'N,
124ø0'W), each of which surpasses the scolopacid longevity record (Marks
1992) and nearly triples the longevity record (8 yr-7 mo) for Marbled
Godwits (Clappet al. 1982).
RESULTS AND DISCUSSION
On 29 Aug. 1992, one of us (MGD) using a 22X spotting scope re-
corded the band number (564-44600) of a godwit, which roosted on pil-
ings at a distance of 20 m. On 2 Dec. 1993, two of us (MAC, OEW) using
a 30X spotting scope recorded the band number (564-44602) of the sec-
ond godwit foraging along tidal channels within 15 m of our observation
point; this godwit has been resighted frequently by students at Humboldt
State University and its band number recorded on at least three subse-
quent occasions by independent observers (MAC; K. Morgan, pers.
Current address: Kings River College, Reedley, California 93654 USA.
2 Current address.' South Carolina Wildlife and Marine Resources Department, Route 2 Box 167,
Greenpond, South Carolina 29446 USA.
TABLE 1. Observations of nine adult Marbled Godwits banded between August 1968 and
November 1969 and subsequently recovered or resighted in Humboldt County, Califor-
nia.
Mini-
Date recovered mum
Bird Date or most recent age
(band 4/) banded resighting (yr-mo) Comnents
564-44541 15 Aug. 1969 ? Jan. 1970 1-5 Unknown cause of death
564-44753 10 Oct. 1969 18 Sep. 1970 1-11 Collected under permit
564-44748 10 Oct. 1969 20 Dec. 1971 3-2 Found dead
564-44722 10 Oct. 1969 15Jan. 1977 8-7 Found dead (Clappet al. 1982)
564-44802 11 Oct. 1969 18 Oct. 1981 12-0 Shot by hunters
564-44600 15 Aug. 1969 29 Aug. 1992 24-2 Observed foraging
564-44602 15 Aug. 1969 14 Mar. 1994 25-9 Observer foraging; limped
noticeably
56444782 15 Aug. 1969 14 Mar. 1994 25-9 Observed foraging
564-44505 10 Oct. 1969 14 Mar. 1994 25-9 Observed foraging
comm.; M. Taft, pers. comm.). On 14 Mar. 1994, MAC observed two ad-
ditional, banded godwits (564-44782; 564-44505) foraging along tidal
channels at a distance of <20 m. All four godwits were banded on the
right tibiotarsus, and we had no difficulty discerning band numbers. Al-
though each of the observers knew of earlier banding efforts at Humboldt
Bay (Gerstenberg 1972), at the time of observation none of us had knowl-
edge of band numbers. Banding schedules (RHG) showed that these four
godwits had been cannon-netted and banded as adults on two separate
occasions (15 August (n = 3); 10 October (n = 1)) in 1969 at the mouth
of Jacoby Creek, Humboldt County, California, within 2 km of the resight-
ing locations. Hence, at the time of most recent observation, the mini-
mum age of the first godwit (564-44602) was 24 yr-2 mo, whereas the
other birds were minimally 25 yr-9 mo old.
Of 256 Marbled Godwits banded by Gerstenberg (1972) at Humboldt
Bay between July 1968 and November 1969, nine have been recovered or
resighted locally (Table 1). The four birds reported here were among 138
banded on 15 Aug. and 10 Oct. 1969. On the basis of the latter sample,
minimum survival probabilities of Marbled Godwits approximate 87%.
From analysis of band recoveries for European shorebirds, Boyd (1962)
estimated survival rates were 56-75%. Boyd's estimates are considerably
lower than minimum annual survival estimates based on resightings of
individually color-marked birds (nearly all of which exceed 70%), prob-
ably because loss of bands has been interpreted as "early mortality" for
some individuals (Evans and Pienkowski 1984). Our observations of Mar-
bled Godwit longevity support estimates based on annual return rates
reported by Evans and Pienkowski (1984). The ability to read numbers
on godwit bands (size 4) in excess of 20 yr old on birds using salt water
habitats is remarkable. Furthermore, it suggests that analyses of North
American band recovery data (for similar bands) may not be plagued by
problems confronting analyses of European banding and recovery re-
cords.
ACKNOWLEDGMENTS
We thank the Department of Wildlife, Humboldt State University for research support. S.
Harris provided banding histories from Humboldt Bay. J. Dunk, S. Landrum and J. Marks
provided helpful comments on the manuscript.
LITERATURE CITED
BOYD, H. 1962. Mortality and fertility of European Charadrii. Ibis 104:368-387.
CLAPP, R. B., M. K. KLIMKIEqCZ, AND J. H. KENNARD. 1982. Longevity records of North
American birds: gaviidae through alcidae. J. Field Ornithol. 53:81-124.
EVANS, P. R., AND M. W. PIENKOWSKI. 1984. Population dynamics of shorebirds. Pp. 83-123,
inJ. Burger and B. L. Olla, eds. Shorebirds: breeding behavior and populations. Plenum
Press, New York, New York.
GERSTENBERG, R. H. 1972. A study of shorebirds (Charadrii) in Humboldt Bay, California--
1968 to 1969. M.Sc. thesis, Humboldt State Univ., Arcata, California.
MARKS, J. s. 1992. Longevity record for the Bristle-thighed Curlew: an extension. J. Field
Ornithol. 63:309-310.
Received 9 Mar. 1994; accepted 6 Jun. 1994.