IN his "Check-List of Birds of the World" James L. Peters (1940: 236)
follows Ridgway and Hellmayr in listing Chaetura vauxi, Chaetura
richmondi, and Chaetura gaumeri as distinct species, in spite of the fact
that other able present-day workers have called them "merely repre-
sentative forms" of the same species (see, in particular, Griscom, 1932:
196). The writer has been obliged to study these three forms carefully
in identifying recent collections from Mexico, and he is now convinced
that they are conspecific.
There is no escaping the fact that they look alike. Their behavior,
nesting habits, and call notes are said to be much the same. They are
all essentially Middle American, even the northward ranging, strongly
migratory vauxi returning to its tropical ancestral home for the winter
months. The ranges of gaumeri and richmondi probably adjoin
in Campeche and Quintana Roo, since no treeless plain, high sierra, or
other barrier appears to exist between them. As for richmondi and
vauxi, or gaumeri and vauxi, though the present known breeding range
of the latter is, in either case, far removed from that of the former, it
is quite conceivable (a) that vauxi, in pushing northward, has only
recently become truly isolated or (b) that further work in the Rocky
Mountains and the Sierra Madre will reveal the fact that a chain of
breeding populations actually connects the three forms. The primary
purpose of the present paper is to announce the discovery of one such
breeding population, a form that is close to gaumeri and between vauxi
and richmondi in color and size.
This new bird has been called "richmondi" by Phillips (1911: 77),
Ridgway (1911: 720), and others. It has also been called "vauxi" by
no less able an ornithologist than Peters who, in examining our material,
perceived it to be darker and smaller than topotypical specimens of
"Cypcelus Vauxi" (type locality Columbia River ----- Fort Vancouver,
Washington) and who may even have suspected it of being a race of the
northwestern United States bird. Need more be said as to its linking
C. v. richmondi and C. v. vauxi? Since no name thus far given to any
Middle American swift appears to be available, the author suggests
Chaetura vauxi tamaulipensis, subsp. nov.
T3/4,E.--Breeding male (testes greatly enlarged) in unworn plumage,
collection of George M. Sutton, No. 9601; Rancho Rinconada, along the
Sabinas River, a mile west of the main Mexico City highway, at an
elevation of about 500 feet, in the vicinity of the village of G6mez
Farias, southwestern Tamaulipas, April 15, 1941; collected by George
Miksch Sutton.
232 THE WILSON BULLETIN December, 1941
Vol. 53, No. 4
SUBSPECIFIC CHARACTERs.--SimiIar to Chaetura vauxi vauxi in
wing-length and body-size, hence larger than C. v. richmondi (from
southern Veracruz, Costa Rica, Guatemala, and interlying parts of
Central America), C. v. gaumeri (from Yucatan), and C. v. ochropygia
(from the Azuero Peninsula of Panama). Darker than C. v. vauxi, es-
pecially on the upper parts, belly, under tail coverts, and basal part of
the throat plumage; crown, back, and wings noticeably more glossy, the
shine being rather strongly greenish; and eyebrow without fine whitish
line that is usually so distinct in C. v. vauxi. Less blackish than C. v.
richmondi above; wings duller, with greenish rather than blue or bluish
green gloss; and rump and under parts somewhat lighter. Less blackish
on back, crown, and wings even than C. v. gaumeri, and more brownish
throughout belly, under tail coverts, and rump. Larger and noticeably
darker-rumped than C. v. ochropygia, a far-removed form that it would
hardly be expected to resemble at all closely.
RAr..--Breeds in heavily wooded lowlands, along streams, in
southwestern Tamaulipas, southeastern San Luis Potosi, and probably
contiguous parts of Hidalgo, northern Veracruz, and southeastern Nuevo
Leon. May be migratory, since one November specimen from Guatemala
has been examined. Probably intergrades with C. v. gaumeri and
C. v. richmondi.
RxARrZs.--There are some misconceptions concerning certain races
of Chaetura vauxi. Two of these have to do with C. v. gaumeri. Through
the courtesy of Dr. Josselyn Van Tyne the author has been able to
examine several freshly plumaged specimens of this race taken recently
at Chichen Itzg, Yucatan. These are, to be sure, less blackish on the
crown and back, and less bluish on the wings than strictly comparable
C. v. richmondi from Costa Rica; but the rump is not strikingly light
by comparison, and there is little evidence in support of calling the race
a pale one. Furthermore, in all freshly plumaged birds the tail is fully
spined. There are several such well-spined individuals in the University
of Michigan Museum of Zoology collection. The spines apparently are
subjected to much abrasion in the limestone wells which the swifts in-
habit, and worn specimens have given rise to the belief that the tail of
C. v. gaumeri is spineless.
A third misconception has to do with certain Guatemalan specimens
that have been called by Griscom (1932: 196) "obvious intermediates"
between C. v. vauxi and C. v. richmondi. The author, who has examined
two of these very birds, feels that the Finca Carolina male, which Gris-
corn describes as "nearer richmondi in size, nearer vauxii in color"
(Amer. Mus. Nat. Hist. No. 393,915) is clearly an example of C. v.
vauxi, and not in any genetic sense of the word intermediate. It is a
molting bird, with badly worn outer primaries, and therefore wholly
unreliable wing-length (111 mm.). The incoming remiges are naturally
dark and glossy, just as they are in freshly plumaged or molting United
George M. NEW RACE OF CHAETURA 233
Sutton
States C. v. vauxl. The San Lucas male, said to be "nearer richmondi
in color, near vauxii in size" (Amer. Mus. Nat. Hist. No. 393,928),
agrees perfectly with the type series of C. v. tamaulipensis and must,
for the present, be considered a Guatemala record of the race; but it
should not be called intermediate between C. v. vauxi and C. v. rich-
mondi, for such a designation implies an actual commingling of the two
forms during the breeding season.
Migrating C. v. vauxi probably move northward and southward to
the west of the breeding grounds of C. v. tamaulipensis, so transient
vauxi-like birds seen in eastern Mexico are likely to be tamaulipensis.
Wintering swifts will have to be identified with great care, however, in
view of Lowery's recent discovery of C. v. vaux in winter in Louisiana
(1939).
MEASUREMENTs.--Type: wing, 111 mm.; tail, 35. Topotypical
male: wing, 111; tail, 37. Topotypical females: wing, 115, 109; tail,
33.5, 32. Female from Matlapa, near Tamazunchale, San Luis Potosi:
wing, 113; tail, 38.5. Female from San Lucas, Guatemala: wing, 114;
tail, 37. Male from Guiaves, Tamaulipas: wing, 113.5; tail, 38. Female
from Carricitos, Tamaulipas: wing, 111; tail, 40. Average of males:
wing, 111.8; tail, 36.6; of females: wing, 112.4; tail, 36.2. Wing meas-
urements are of the chord of the folded manus.
ACNOWLEDGEMENTs.--The valuable assistance of Mr. James L.
Peters and Dr. Josselyn Van Tyne has already been mentioned. Mr.
Adriaan J. van Rossem was good enough to lend the author a useful
series of C. v. vauxi (including virtual topotypes from western Wash-
ington) from the Dickey Collections. The American Museum of Nat-
ural History cooperated by lending Guatemalan and Costa Rican ma-
terial. Mr. George H. Lowery, Jr., was particularly helpful in lending
all of his freshly plumaged, carefully washed C. v. vauxi taken recently
at Baton Rouge, Louisiana.
REFERENCES CITED
GRISCO/f, LUDLOW
1932 The Distribution of BirdsLife in Guatemala. Bull. Amer. Mus. Nat.
Hist., 64.
LOWERY, GEORCE H., JR.
1939 Vaux's Swift in Louisiana. Wilson Bulletin, 51: 199-201.
PETERS, JAMES L.
1940 Check-list of Birds of the World, Vol. 4. Harvard University Press,
Cambridge.
PaILLIPS, Joas C.
1911 A Year's Collecting in the State of Tamaulipas, Mexico. Auk, 28:
67-89.
RIDGWA3/4, ROBERT
1911 The Birds of North and Middle America. Bull. U.S. Nat. Mus., No. 50,
part 5.
CORNELL UNIVERSITY, ITHACA, NEW YORK.